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- (-) Remove Animalia- filter Animalia-
- (-) Remove Arthropods- filter Arthropods-
- Animals- (2) Apply Animals- filter
- Arthropoda- (2) Apply Arthropoda- filter
- Behavior- (1) Apply Behavior- filter
- Coleoptera-: Insecta- (2) Apply Coleoptera-: Insecta- filter
- dung-rolling (1) Apply dung-rolling filter
- Insects- (2) Apply Insects- filter
- Invertebrata- (2) Apply Invertebrata- filter
- Invertebrates- (2) Apply Invertebrates- filter
- male- (2) Apply male- filter
- male-dimorphism (1) Apply male-dimorphism filter
- male-participation (1) Apply male-participation filter
- mating-effort (1) Apply mating-effort filter
- morphology- (1) Apply morphology- filter
- nes (1) Apply nes filter
- nest-building (1) Apply nest-building filter
- Onthophagus-taurus [horned-beetle] (Coleoptera-): female- (1) Apply Onthophagus-taurus [horned-beetle] (Coleoptera-): female- filter
- parental-care-patter (1) Apply parental-care-patter filter
- Reproduction- (1) Apply Reproduction- filter
- Scarabaeus-catenatus [dung-beetle] (Coleoptera-): female- (1) Apply Scarabaeus-catenatus [dung-beetle] (Coleoptera-): female- filter