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- (-) Remove Animalia- filter Animalia-
- (-) Remove mating-effort filter mating-effort
- (-) Remove nest-building filter nest-building
- Animals- (1) Apply Animals- filter
- Arthropoda- (1) Apply Arthropoda- filter
- Arthropods- (1) Apply Arthropods- filter
- Behavior- (1) Apply Behavior- filter
- Coleoptera-: Insecta- (1) Apply Coleoptera-: Insecta- filter
- dung-rolling (1) Apply dung-rolling filter
- Insects- (1) Apply Insects- filter
- Invertebrata- (1) Apply Invertebrata- filter
- Invertebrates- (1) Apply Invertebrates- filter
- male- (1) Apply male- filter
- male-participation (1) Apply male-participation filter
- nes (1) Apply nes filter
- Scarabaeus-catenatus [dung-beetle] (Coleoptera-): female- (1) Apply Scarabaeus-catenatus [dung-beetle] (Coleoptera-): female- filter