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- (-) Remove Animalia- filter Animalia-
- (-) Remove Invertebrata- filter Invertebrata-
- (-) Remove male-participation filter male-participation
- Animals- (1) Apply Animals- filter
- Arthropoda- (1) Apply Arthropoda- filter
- Arthropods- (1) Apply Arthropods- filter
- Behavior- (1) Apply Behavior- filter
- Coleoptera-: Insecta- (1) Apply Coleoptera-: Insecta- filter
- dung-rolling (1) Apply dung-rolling filter
- Insects- (1) Apply Insects- filter
- Invertebrates- (1) Apply Invertebrates- filter
- male- (1) Apply male- filter
- mating-effort (1) Apply mating-effort filter
- nes (1) Apply nes filter
- nest-building (1) Apply nest-building filter
- Scarabaeus-catenatus [dung-beetle] (Coleoptera-): female- (1) Apply Scarabaeus-catenatus [dung-beetle] (Coleoptera-): female- filter