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- (-) Remove dung-beetle (Coleoptera-) filter dung-beetle (Coleoptera-)
- (-) Remove Iberian-Peninsula (Europe- filter Iberian-Peninsula (Europe-
- (-) Remove Population-Studies filter Population-Studies
- Animalia- (1) Apply Animalia- filter
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- Coleoptera-: Insecta- (1) Apply Coleoptera-: Insecta- filter
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- Scarabaeidae- (Coleoptera-) (1) Apply Scarabaeidae- (Coleoptera-) filter