02147nas a2200265   4500008004100000245012400041210006900165300001000234490000700244520128800251653001101539653001201550653001701562653001101579653001701590653001401607653001401621653001601635653001701651653001501668653001401683100002701697700001901724856013801743       2003                        eng d00aPhylogeny of Scarabaeus (Pachysoma MacLeay) stat. nov., and related flightless Scarabaeini (Scarabaeidae: Scarabaeinae)0 aPhylogeny of Scarabaeus Pachysoma MacLeay stat nov and related f  a47-710 v403 aThemonophylyofPachysomaMacLeayisassessedviacladisticanalysis,basedonsixty-fouradultcharactersfrom thirty-seventaxacomprisingall currentlyandpreviously recognizedgenerainScarabaeini, except Madateuchus Paulian.PachysomacomprisesamonophyleticcladeofderivedspeciesofScarabaeusandNeopachysomaFerreira isthesisterlineageofPachysomasensustricto.Scarabaeus(Pachysoma)stat. nov. isconsideredasubgenusof ScarabaeusLinnaeuswhileNeopachysomaFerreiraismaintainedasasynonymof Pachysomasensulato. NeomnematiumJanssenstogether withMnematiumMacLeayispolyphyletic, whilethelatter isparaphyletic. MnematiumMacLeayandNeomnematiumJanssensaremaintainedassynonymsofScarabaeus,andtheirspecies includedinsubgeneraScarabaeus(Scarabaeus) andScarabaeus(Scarabaeolus).DrepanopodusJanssens, KheperJanssensandSceliagesWestwood,asderivedcladesofScarabaeus,couldalsoberecognizedassubgen- eratomaintainamonophyleticScarabaeus,but aremaintainedasgenerapendingfurther work. Aridificationof theNamibDesertispostulatedtohaveinitiatedtheevolutionofScarabaeus(Pachysoma),whichfeedondrydungor detritusandarederivedfromaball-rollingandwet-dung-feedingScarabaeus-likeancestor. Theevolutionof flightlessnessandfoodrelocationstrategy,biogeography,andadditionaldistributionalandbiologicalinformationare presented for the group. 10aAptery10aBiology10aDrepanopodus10aKheper10aPachylomerus10aPachysoma10aphylogeny10aPsammophily10aScarabaeolus10aScarabaeus10aSceliages1 aHarrison, James, du G.1 aPhilips, Keith  uhttps://scarabnet.myspecies.info/phylogeny-scarabaeus-pachysoma-macleay-stat-nov-and-related-flightless-scarabaeini-scarabaeidae-scar