G. M.  Carpaneto, Mazziotta, A., and Piattella, E., ?Changes in food resources and conservation of scarab beetles: from sheep to dog dung in a green urban area of Rome (Coleoptera, Scarabaeoidea)?, Biological Conservation, vol. 123, pp. 547-556, 2005.

A. L. V.  Davis, ?Community organization in a South African, winter rainfall, dung beetle assemblage (Coleoptera: Scarabaeidae)?, Acta Oecologica, vol. 15, pp. 727-738, 1994.

A.  Gonzalez-Megias and Sanchez-Pinero, F., ?Effects of brood parasitism on host reproductive success: evidence from larval interactions among dung beetles?, Oecologia, vol. 134, pp. 195-202, 2003.

P.  Jay-Robert, Niogret, J., Errouissi, F., Labarussias, M., Paoletti, E., Luis, M. Vazques, and Lumaret, J. - P., ?Relative efficiency of extensive grazing vs. wild ungulates management for dung beetle conservation in a heterogeneous landscape from Southern Europe (Scarabaeinae, Aphodiinae, Geotrupinae)?, Biological Conservation, vol. 141, pp. 2879-2887, 2008.

J. A.  Finn and Giller, P. S., ?Experimental investigations of colonisation by north temperate dung beetles of different types of domestic herbivore dung?, Applied Soil Ecology, vol. 20, pp. 1-13, 2002.

T.  Roslin and Koivunen, A., ?Distribution and abundance of dung beetles in fragmented landscapes?, Oecologia, vol. 127, pp. 69-77, 2001.