C. M.  House and Simmons, L. W., ?Genital morphology and fertilization success in the dung beetle Onthophagus taurus: an example of sexually selected male genitalia?, Proceedings of the Royal society of London B, vol. 270, pp. 447-455, 2003.

C. M.  House and Simmons, L. W., ?The evolution of male genitalia: patterns of genetic variation and covariation in the genital sclerites of the dung beetle Onthophagus taurus?, Journal of Evolutionary Biology, vol. 18, pp. 1281-1292, 2005.

C. M.  House and Simmons, L. W., ?Offensive and defensive sperm competition roles in the dung beetle Onthophagus taurus (Coleoptera: Scarabaeidea)?, Behavioral Ecology and Sociobiology, vol. 60, pp. 131-136, 2006.

J.  Hunt and Simmons, L. W., ?Patterns of fluctuating asymmetry in beetle horns: an experimental examination of the honest signalling hypothesis?, Behavioral Ecology and Sociobiology, vol. 41, pp. 109-114, 1997.

J.  Hunt and Simmons, L. W., ?Patterns of parental provisioning covary with male morphology in a horned beetle (Onthophagus taurus) (Coleoptera: Scarabaeidae)?, Behavioral Ecology and Sociobiology, vol. 42, pp. 447-451, 1998.

J.  Hunt and Simmons, L. W., ?Confidence of paternity and paternal care: covariation revealed through the experimental manipulation of the mating system in the beetle Onthophagus taurus?, Journal of Evolutionary Biology, vol. 15, pp. 784-795, 2002.

J.  Hunt and Simmons, L. W., ?The genetics of maternal care: direct and indirect genetic effects on phenotype in the dung beetle Onthophagus taurus?, Proceedings of the National Academy of Sciences, vol. 99, pp. 6828-6832, 2002.

J.  Hunt and Simmons, L. W., ?Optimal maternal investment in the dung beetle Onthophagus taurus??, Behavioral Ecology and Sociobiology, vol. 55, pp. 302-312, 2004.

J. S.  Kotiaho and Simmons, L. W., ?Effects of Macrocheles mites on longevity of males of the dimorphic dung beetle Onthophagus binodis?, Journal of Zoology, vol. 254, pp. 441-445, 2001.

A. P.  Moczek, Hunt, J., Emlen, D. J., and Simmons, L. W., ?Threshold evolution in exotic populations of a polyphenic beetle?, Evolutionary Ecology Research, vol. 4, pp. 587-601, 2002.