M.  Bertone, ?Dung beetles (Coleoptera: Scarabaeidae and Geotrupidae) in North Carolina Pasture Ecosystem?, North Carolina University, Raleigh, 2006.

M. A.  Bertone, ?Dung beetles (Coleoptera: Scarabaeidae and Geotrupidae) of North Carolina Cattle Pastures  and their Implications for Pasture Improvement?, North Carolina University, Raleigh, 2004.