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A. J.  Davis, ?The ecology and behaviour of rainforest dung beetles in northern Borneo.?, University of Leeds, Leeds, 1993.

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I. C.  Mittal, ?Natural Manuring and Soil Conditioning by Dung Beetles?, Tropical Ecology, vol. 34, pp. 150-159, 1993.

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K.  Vulinec, ?A southern range extension for the introduced dung beetle Onthophagus taurus Schreber (Coleoptera: Scarabaeidae)?, The Coleopterists Bulletin, vol. 47, pp. 129-130, 1993.

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