J. S.  Kotiaho and Simmons, L. W., ?Effects of Macrocheles mites on longevity of males of the dimorphic dung beetle Onthophagus binodis?, Journal of Zoology, vol. 254, pp. 441-445, 2001.

C.  Palestrini, Barbero, E., and Rolando, A., ?Intra- and interspecific aggregation among dung beetles (Coleoptera: Scarabaeoidea) in an Alpine pasture?, Journal of Zoology, vol. 245, pp. 101-109, 1998.

C.  Palestrini and Rolando, A., ?Body size and paternal invest ment in the genus Onthophagus (Coleoptera, Scarabaeoidea)?, Journal of Zoology, vol. 255, pp. 405-412, 2001.