C.  Avendano-Mendoza, Moron-Rios, A., Cano, E. B., and Leon-Cortes, J. L., ?Dung beetle community (Coleoptera : Scarabaeidae : Scarabaeinae) in a tropical landscape at the Lachua Region, Guatemala?, Biodiversity and Conservation, vol. 14, pp. 801-822, 2005.