Y.  Cambefort, ?Body-Size, Abundance, and Geographical-Distribution of Afrotropical Dung Beetles (Coleoptera, Scarabaeidae)?, Acta Oecologica, vol. 15, pp. 165-179, 1994.

A. L. V.  Davis, ?Community organization in a South African, winter rainfall, dung beetle assemblage (Coleoptera: Scarabaeidae)?, Acta Oecologica, vol. 15, pp. 727-738, 1994.