A. L. V.  Davis, ?Diel and seasonal community dynamics in an assemblage of coprophagous, Afrotropical, dung beetles (Coleoptera: Scarabaeidae s. str., Aphodiidae, and Staphylinidae: Oxytelinae)?, Journal of African Zoology, vol. 110, pp. 290-308, 1996.

A. L. V.  Davis, ?Dung beetle diversity in South Africa: influential factors, conservation status, data inadequacies and survey design?, African Entomology, vol. 10, pp. 53-65, 2002.

A.  Gonzalez-Megias and Sanchez-Pinero, F., ?Effects of brood parasitism on host reproductive success: evidence from larval interactions among dung beetles?, Oecologia, vol. 134, pp. 195-202, 2003.

T.  Roslin and Koivunen, A., ?Distribution and abundance of dung beetles in fragmented landscapes?, Oecologia, vol. 127, pp. 69-77, 2001.