P.  Hirschberger and Degro, H., ?Oviposition of the dung beetle Aphodius ater in relation to the abundance of yellow dungfly larvae (Scataphaga sterocoraria)?, Ecological Entomology, vol. 21, pp. 352-357, 1996.

S. A.  Hutton and Giller, P. S., ?Intra- and interspecific aggregation of north temperate dung beetles on standardised and natural dung pads: the influence of spatial scale?, Ecological Entomology, vol. 29, pp. 594-605, 2004.

P.  Holter and Scholtz, C. H., ?Are ball-rolling (Scarabaeini, Gymnopleurini, Sisyphini) and tunnelling scarabaeine dung beetles equally choosy about the size of ingested dung particles??, Ecological Entomology, vol. 30, pp. 700-705, 2005.

K.  Vessby, ?Habitat and weather affect reproduction and size of the dung beetle Aphodius fossor?, Ecological Entomology, vol. 26, pp. 430-435, 2001.