H.  Carrillo-Ruiz and Moron, M. - A., ?Fauna de Coleoptera Scarabaeoidea de Cuetzalan del Progreso, Puebla, Mexico?, Acta Zoologica Mexicana (nueva serie), vol. 88, pp. 87-121, 2003.

A. L. V.  Davis, ?Dung beetle diversity in South Africa: influential factors, conservation status, data inadequacies and survey design?, African Entomology, vol. 10, pp. 53-65, 2002.

J. M.  Lobo and Martin-Piera, F., ?Between-group differences in the Iberian dung beetle species-area relationship (Coleoptera : Scarabaeidae)?, Acta Oecologica-International Journal of Ecology, vol. 20, pp. 587-597, 1999.

C. A.  Medina, S., F. Escobar, and Kattan, G. H., ?Diversity and habitat use of dung beetles in a restored Andean landscape?, Biotropica, vol. 34, pp. 181-187, 2002.

J.  Noriega, Realpe, E., and Fagua, G., ?Diversidad de escarabajos coprofagos (Coleoptera: Scarabaeidae) en un bosque de galeria con tres estadios de alteracion?, Revista de la Facultad de Ciencias, Univ. Javeriana, Bogota, vol. 12, pp. 51-63, 2007.

L. Astrid Pul Herrera, Canas, R. Antonio Ri, Harders, F. Gast, and von Hildebrand, P., ?Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) del Parque Nacional Natural "Serrania de Chiribiquete", Caqueta, Colombia (Parte I)?, m3m: Monografias Tercer Milenio, vol. 3, pp. 51-58, 2003.

D. C.  Ronqui and Lopes, J., ?Composicao e diversidade de Scarabaeoidea (Coleoptera) atraidos por armadilha de luz em area rural no norte do Parana, Brasil?, Iheringia, Servicio Zoologico do Porto Alegre, vol. 96, pp. 103-108, 2006.

Y.  Xiao-Dong, Tian-Hong, L., and Hong-Zhang, Z., ?Species composition and diversity of scarab beetle (Coleoptera: Scarabaeoidea) communities at Dongling Mountain, Beijing?, Biodiversity Science, vol. 11, pp. 179-187, 2003.