| Publication Type: | Journal Article |
| Year of Publication: | 1997 |
| Authors: | D. J. Emlen |
| Journal: | Behavioral Ecology and Sociobiology |
| Volume: | 41 |
| Pagination: | 335-341 |
| Date Published: | Aug |
| Keywords: | alternative reproductive behavior, horned beetles, male competition, male dimorphism, Onthophagus |