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- (-) Remove Animals- filter Animals-
- (-) Remove Coleoptera-: Insecta- filter Coleoptera-: Insecta-
- (-) Remove horn-: size- filter horn-: size-
- Animalia- (1) Apply Animalia- filter
- Arthropoda- (1) Apply Arthropoda- filter
- Arthropods- (1) Apply Arthropods- filter
- Behavior- (1) Apply Behavior- filter
- body-size (1) Apply body-size filter
- brood-mass (1) Apply brood-mass filter
- Evolution-and-Adaptation (1) Apply Evolution-and-Adaptation filter
- Insects- (1) Apply Insects- filter
- Invertebrata- (1) Apply Invertebrata- filter
- Invertebrates- (1) Apply Invertebrates- filter
- male- (1) Apply male- filter
- male-p (1) Apply male-p filter
- Onthophagus-taurus [dung-beetle] (Coleoptera-): female- (1) Apply Onthophagus-taurus [dung-beetle] (Coleoptera-): female- filter