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- (-) Remove Coleoptera-: Insecta- filter Coleoptera-: Insecta-
- (-) Remove dung-beetles filter dung-beetles
- Animalia- (1) Apply Animalia- filter
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- ecological-analysis: analytical-method (1) Apply ecological-analysis: analytical-method filter
- human (1) Apply human filter
- Insects- (1) Apply Insects- filter
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- Invertebrates- (1) Apply Invertebrates- filter
- Methods-and-Techniques (1) Apply Methods-and-Techniques filter
- Scarabaeinae- (Coleoptera-): indicator-group (1) Apply Scarabaeinae- (Coleoptera-): indicator-group filter