@article {1530,
	title = {Trophic preferences mediated by olfactory cues in dung beetles colonizing cattle and horse dung},
	journal = {Environmental Entomology},
	volume = {33},
	number = {2},
	year = {2004},
	note = {digital copy},
	month = {Apr},
	pages = {370-377},
	abstract = {Scarab beetles colonizing vertebrate feces are considered polyphagous insects, because they often use dung of several species. However, recent works have provided evidence for strong feeding preferences in some dung beetle species. The influence of dung odors in the process of resource selection by beetles has not received attention. The objective of this study was to investigate trophic preferences in Mediterranean dung beetles colonizing cattle and horse dung. The pattern of resource use by insects was studied using series of pitfall traps baited with cattle or horse dung in three different sites of south-central France. A total of 4,276 insects belonging to 39 species were captured. Cattle dung attracted more insects than horse feces (2,570 versus 1,706 individuals). Twenty-four of the 39 beetle species collected had clear feeding preferences for cattle dung (13 species) or horse dung (11 species). None of these species seemed linked exclusively to one kind of dung. The behavioral responses of seven scarab beetles to volatile compounds emitted by cattle and horse dung were compared in laboratory olfactometer bioassays. Three insect species (Aphodius erraticus, A. scrutator, Onthophagus vacca) were more attracted to volatile compounds from cattle dung. Two other species (Euonthophagus amyntas, Bubas bubalus) orientated preferentially toward horse dung volatiles. Except for two species, beetles were thus attracted to volatiles from the dung type they preferred in the field. The reasons why coprophagous beetles show local feeding preferences for particular dung types are discussed.},
	keywords = {bait, burial, capacity, cattle dung, COLEOPTERA, colonization, communities, dung beetle, Dung beetles, feces, food, horse dung, mammal mammals, olfactometer, Scarabaeidae, Scarabaeoidea, trophic preferences},
	url = {<Go to ISI>://000220856500033},
	author = {Dormont, L. and Epinat, G. and Lumaret, J. P.}
}
@article {1926,
	title = {Convergences and divergences between two European mountain dung beetle assemblages (Coleoptera, Scarabaeoidea)},
	journal = {Animal Biodiversity and Conservation},
	volume = {30},
	number = {1},
	year = {2007},
	pages = {83-96},
	abstract = {We analyzed the altitudinal change in dung beetle species richness and the relative proportion of higher taxa, as well as the turnover in the type of distribution and range size of species in two mountain chains located at the two extremes of Europe (Western Rhodopes Mountains and the Iberian Central System). Both mountain ranges showed a clear substitution among higher taxa (Aphodiinae{\textendash}Geotrupinae vs. Scarabaeidae) and species richness variation with the altitude was similar. We suggest that East European dung beetle assemblages are conditioned by a horizontal colonization process in which mountains had been reached in relatively recent geological time by elements coming from different latitudes. In spite of these convergences, Rhodopes dung beetle assemblages are characterized by a significantly lower proportion of narrowly distributed species and a lower relevance of Aphodiinae species in lowland places. Although these divergences can be partially attributed to the dissimilar sampling effort accomplished in both regions, we suggest that the low number on narrowly distributed species could be due to the different role of these two mountain zones as refuges during glaciar{\textendash}interglaciar Pleistocene cycles.},
	keywords = {Altitudinal variation, Dung beetles, Iberian Central System, Refuges, Rhodopes mountain range, Scarabaeoidea},
	isbn = {1578-665X},
	author = {Lobo, Jorge M. and Gueorguiev, B. and Chehlarov, E.}
}
@article {1932,
	title = {Diversity, distinctiveness and conservation status of the Mediterranean coastal dung beetle assemblage in the Regional Natural Park of the Camargue (France)},
	journal = {Diversity and Distributions},
	volume = {7},
	year = {2001},
	pages = {257-270},
	abstract = {The Mediterranean region as a whole has the highest dung beetle species richness within Europe. Natural coastal habitats in this region are among those which have suffered severe human disturbance. We studied dung beetle divers- ity and distinctiveness within one of the most important coastal protected areas in the west Euro- Mediterranean region (the regional Park of Camargue, southern France) and made comparisons of dung beetle assemblages with other nearby Mediterra- nean localities, as well as with other coastal pro- tected area (Doñana National Park, Spain). Our finding showed that: (1) The species richness of coastal habitats in the Camargue is low and only grasslands showed a similar level of species rich- ness and abundance to inland habitats of other Mediterranean localities. The unique habitats of the coastal area (beaches, dunes and marshes) are largely colonized by species widely distributed in the hinterland. (2) In spite of their low general distinctiveness, dune and marsh edges are charac- terized by the occurrence of two rare, vulnerable, specialized and large roller dung beetle species of the genus Scarabaeus. As with other Mediterranean localities, current findings suggest a recent decline of Scarabaeus  populations and the general loss of coastal dung beetle communities in Camargue. (3) The comparison of dung beetle assemblages between the Camargue and Doñana shows that, in spite of the low local dung beetle species rich- ness in the Camargue, the regional dung beetle diversity is similar between both protected areas. Unique historical and geographical factors can explain the convergence in regional diversity as well as the striking divergence in the composition of dung beetle assemblages between both territories.},
	keywords = {Don, Dung beetles, insect coastal conservation, Mediterranean region, Scarabaeoidea, the Camargue, ̃ana},
	author = {Lobo, Jorge M. and Lumaret, Jean-Pierre and Jay-Robert, Pierre}
}
@article {1955,
	title = {Geographic distribution of endemic dung beetles (Coleoptera, Scarabaeoidea) in the Western Palaeartic region},
	journal = {Biodiversity Letters},
	volume = {3},
	number = {6},
	year = {1996},
	pages = {192-199},
	abstract = {The distribution of endemic dung beetle species has been analysed in the West Palaearctic region (547spp.). These species are concentrated in the Mediterranean Basin or in the neighbouring regions, with two main centres of endemism located on opposite sides of thls Basin. The centres of endemism and the degree of endemicity were very different according to the three main dung beetle families. The possible explanation of these patterns has been examined.},
	keywords = {centres of endemism, COLEOPTERA, Dung beetles, endemic species distributions., Mediterranean region, Scarabaeoidea},
	author = {Lumaret, Jean-Pierre and Lobo, Jorge M.}
}
@article {2277,
	title = {Larval morphology and breeding behavior of the genus Pedaridium Harold (Coleoptera: Scarabaeidae)},
	journal = {Annals of the Entomological Society of America},
	volume = {94},
	number = {4},
	year = {2001},
	pages = {596-604},
	abstract = {The third-instar larva and breeding behavior of Pedaridium almeidai Pereira and P. brasiliensis Ferreira \& Galileo are described using material collected at Sa ̃ o Paulo, Brazil, and, the third-instar larva of P. adisi (Ratcliffe) is redescribed. The most important morphological charac- teristics for larval stages are found in the epipharynx, the setation of the venter of the last abdominal segment, the endoskeletal lines of the anal lobes, the mandibles, and the respiratory plate of the thoracic spiracle. A key to the known larvae of the genus Pedaridium is also provided. Results show that P. almeidai is more similar to P. adisi than P. brasiliensis. The presence/absence of stridulatory areas on the mandibles, the form of the frontal suture, the form of the sensorial area of the antennae, the form of the molar and scissorial areas of the mandibles, and the different size and form of the lacinia suggest a possible polyphyly of the genus Pedaridium. Results con{\TH}rm the endocoprid breeding behavior of Pedaridium, which together with Trichillum, represents the most primitive breeding pattern of Scarabaeinae. In the species studied, there is neither food relocation nor brood ball construction. This primitive breeding behavior implies exploitation without modi{\TH}cation or relocation of the resource. Possible kleptoparasitism was observed in P. brasiliensis.},
	keywords = {Ateuchini, breeding patterns, Dung beetles, larval description, Scarabaeoidea, taxonomy},
	author = {Verdu, Jose R. and Galante, Eduardo}
}
