@article {1307,
	title = {Patrones de distribucion de escarabajos coprofagos (Coleoptera: Scarabaeidae) en relicto del bosque altoandino cordillera oriental de Colombia},
	journal = {Caldasia},
	volume = {19},
	number = {1-2},
	year = {1997},
	note = {have copy},
	pages = {191-204},
	keywords = {Andes, Colombia, dung beetle, forest relicts, fragmentation (biological conservation 2007), Scarabaeidae},
	author = {Amat-Garcia, German and Lopera Toro, Alejandro and Amezquita, S. J. A.}
}
@article {1309,
	title = {Comparaci{\'o}n de la composici{\'o}n y riqueza de especies de escarabajos copr{\'o}fagos (Coleoptera: Scarabaeidae) en remanentes de bosque de la Orinoquia Colombiana},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {76},
	year = {1999},
	note = {have copy},
	pages = {113-126},
	keywords = {Colombia, community, deforestation, dung beetle (biological conservation 2007), fragmentation, richness, Scarabaeinae},
	author = {Amezquita, Sandra J. and Forsyth, Adrian and Lopera, Alejandro and Camacho, Andr{\'e}s}
}
@article {1405,
	title = {Preferencia por cebo de los escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de un remanente de bosque seco tropical al norte de Tolima (Colombia)},
	journal = {m3m: Monografias Tercer Milenio},
	volume = {3},
	year = {2003},
	pages = {59-65},
	abstract = {Feeding and nesting tactics may limit the effects of species overlapping on one particular resource in a diverse community (Halffter \& Edmonds, 1982). Diversity of feeding habits characterizes the dung beetle groups of the Neotropics, and undoubtedly contributes to to the high species richness of this region (Gill, 1991). Feeding habits determine some ecological aspects (Halffter, 1959) of scarabaeids, like behavior, distribution, morphology and development (Halffter \& Matthews, 1966). This paper studies scarabaeine bait preferences in a dry forest patch and its surrounding matrix. 22 species were captured, with 6 new records for the area. The bait with the highest preference rating was omnivore dung, followed by carrion, fungi, cow dung and rotting fruit. The species with the highest abundance was Dichotomius ca. centralis, with 758  individuals and a copro-necrophagous diet. Its dominance was not restricted to the forested area, and significant numbers were found at the pasture matrix. Strict diet species like Deltochilum valgum agropyge were also collected. The dung beetle community observed was mostly generalist (71\%), possibly due to the scarcity of excrement, which forces individuals to use other sources of food with a similar nutritional value.},
	keywords = {Colombia, diet, dry forest, preferences, resources, Scarabaeinae, Tolima},
	author = {Bustos-Gomez Lida, Fernanda and Lopera Toro, Alejandro}
}
@article {1581,
	title = {Diversidad de Coleopteros coprofagos (Scarabaeidae: Scarabaeinae) en un mosaico de habitats en la reserva natural Nunak, Guaviare, Colombia},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {79},
	year = {2000},
	pages = {103-121},
	keywords = {Amazon, Colombia, diversity, dung beetle, Scarabaeinae, spatial heterogeneity},
	author = {Escobar S. , Federico}
}
@mastersthesis {1941,
	title = {Distribuici{\'o}n y diversidad de escarabajos copr{\'o}fagos (Scarabaeidae: Coleoptera) en tres relictos de bosque altoandino (Cordillera Oriental, Vertiente Occidental), Colombia.},
	year = {1996},
	note = {need copy},
	pages = {123},
	school = {Pontificia Universidad Javeriana},
	type = {Tesis de Grado},
	address = {Bogot{\'a}. Colombia.},
	keywords = {Colombia, Disturbance, dung beetle, fragmentation, moist tropical forest, scarabaeinae (biological conservation 2007)},
	author = {Lopera, A.}
}
@article {2004,
	title = {Diversity and habitat use of dung beetles in a restored Andean landscape},
	journal = {Biotropica},
	volume = {34},
	number = {1},
	year = {2002},
	pages = {181-187},
	keywords = {Andes, Colombia, diversity, dung beetle, habitat use, landscape, restoration, Scarabaeidae},
	author = {Medina, Claudia A. and Escobar S. , Federico and Kattan, Gustavo H.}
}
@article {2006,
	title = {Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia},
	journal = {Biota Colombiana},
	volume = {2},
	number = {2},
	year = {2001},
	pages = {131-144},
	keywords = {COLEOPTERA, Colombia, coprophagous beetles, list of species, Scarabaeidae},
	author = {Medina, Claudia A. and Lopera Toro, Alejandro and Vitolo, Adriana and Gill, Bruce}
}
@article {2079,
	title = {First report of whip scorpions (Arachnida: Uropygi) feeding on dung beetles (Coleoptera: Scarabaeidae)},
	journal = {Boletin de la Sociedad Entomologica Aragonesa},
	volume = {42},
	year = {2008},
	pages = {451-452},
	abstract = {Dung beetles are among the most inrportanl insect groups in ecosystems, but despite th{\textexclamdown}s l{\textexclamdown}ttle information exists about their predators. In the present study, predation on dung beetles by whip scorpions is reported for the first time, based on observations carried out under captivity. Further research and analyses of tl-re predators of this groLrp are necessary.},
	keywords = {Arachnida, COLEOPTERA, Colombia, Dichotomius, Mastigoproctus, predation, Scarabaeidae, Uropygi, whip scorpions},
	author = {Noriega, Jorge and Botero Trujillo, Ricardo}
}
@article {2081,
	title = {Brief note: First report of the genus Tetramereia Klages, 1907 (Coleoptera: Scarabaeidae: Phanaeini) in Colombia - Notes to its distribution},
	journal = {Biota Colombiana},
	volume = {9},
	number = {1},
	year = {2008},
	pages = {133-135},
	keywords = {Colombia, Dung beetles, Phanaeini, Scarabaeidae, Tetramereia},
	author = {Noriega, Jorge and Rengifo, Juan Manuel and Vaz-de-Mello, Fernando Z.}
}
@article {2093,
	title = {Biogeography of the areas and Canthonini (Coleoptera: Scarabaeidae) of dry tropical foress in Mesoamerica and Colombia},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {23},
	number = {1},
	year = {2007},
	pages = {73-108},
	abstract = {This biogeographical analysis examines the historical, geological, climatic and ecological processes that have influenced the formation of the dry tropical forests (DTF) of Mesoamerica and Colombia, areas that are the setting for multiple biogeographical stories that in this case are illustrated by the patterns and evolutionary processes of Canthonini. In this study we test the hypothesis that the Canthonini fauna of dry tropical forests has a South American affinity. To this end, we compare extant species from a tract of dry tropical forest in Mexico, a second enclave in Costa Rica, four from the Caribbean region of Colombia and finally one from the north of Tolima in the Upper Magdalena River Valley, Colombia. The geomorphological characteristics of the enclaves of DTF are also compared, as are the geographical distribution and taxonomic affinities of each of the species found in these dry tropical forests. The biogeographical, historical and geological aspects of the enclaves were evaluated using a Parsimony Analysis of Endemicity (PAE), with two tropical rain forests as the outgroup: Leticia (Amazonas, Colombia) and Los Tuxtlas (Veracruz, Mexico). This study reveals the origin and distribution of Neotropical dry forests in the Pleistocene, and the establishment of its dry conditions during the Holocene. It also reveals apparent similarities among the Canthonini of the dry tropical forests of Mexico, Costa Rica and Colombia, with three geographical distribution patterns that correspond to different degrees of expansion towards the north and the diversification of evolutionary lines, and even species of South American origin. The comparison of the cladogram generated for species of Canthonini with that of the geological events that have occurred in the study regions indicates that the distribution of Canthonini in dry tropical forests began during the Pliocene with the re-establishment of the Panamanian connection, with no evidence of previous geomorphological events having any influence. On the other hand, there are few species shared with the tropical rain forests used as the outgroup for the cladograms. },
	keywords = {biogeographical patterns, Canthonini, Colombia, Costa Rica, dry tropical forest, Mexico, Scarabaeinae},
	author = {Padilla-Gil, Dora Nancy and Halffter, Gonzalo}
}
