@article {1363,
	title = {Molecular phylogeny and biogeography of the dung beetle genus Temnoplectron Westwood (Scarabaeidae : Scarabaeinae) from Australia{\textquoteright}s wet tropics},
	journal = {Molecular Phylogenetics and Evolution},
	volume = {31},
	number = {2},
	year = {2004},
	note = {PDF},
	month = {May},
	pages = {741-753},
	abstract = {The landscape of the Australian Wet Tropics can be described as "islands" of montane rainforest Surrounded by warmer or more xeric habitats. Historical glaciation cycles have caused expansion and contraction of these rainforest "islands" leading to consistent patterns of genetic divergence within species of vertebrates. To explore whether this dynamic history has promoted speciation in endemic and diverse groups Of insects, we used a combination of mtDNA sequencing and morphological characters to estimate relationships and the tempo of divergence among Australian representatives of the dung beetle genus Temnoplectron. This phylogenetic hypothesis shares a number of well-supported clades with a previously published phylogenetic hypothesis based on morphological data. though statistical support for several nodes is weak. Sister species relationships well-supported in both tree topologies. and a tree obtained by combining the two data sets. suggest that speciation has mostly been allopatric. We identify a number of speciation barriers, which coincide with phylogeographic breaks found in vertebrate species. Large sequence divergences between species emphasize that speciation events are ancient (pre- Pleistocene). The flightless, rainforest species appear to have speciated rapidly. but also in the distant past.},
	keywords = {biogeography, Dung beetles, molecular phylogeny, speciation, tropical rainforest},
	url = {<Go to ISI>://000220948000025},
	author = {Bell, K. L. and Yeates, D. K. and Moritz, C. and Monteith, G. B.}
}
@article {1393,
	title = {Effect of vagility potential on dispersal and speciation in rainforest insects},
	journal = {Journal of Evolutionary Biology},
	volume = {17},
	number = {5},
	year = {2004},
	pages = {994-1006},
	keywords = {biogeography, dispersal, Dung beetles, evolution, rainforest, speciation, vagility, vicariance},
	author = {Bouchard, P. and Brooks, D. R.}
}
@article {1447,
	title = {Dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Lanyu},
	journal = {Coll. and Res.},
	volume = {15},
	year = {2002},
	month = {Dec},
	pages = {53-62},
	keywords = {biogeography, dung beetle, Lanyu, new records, Scarabaeinae, Taiwan},
	author = {Chen, Keh-Miin and Yang, Man-Miao and Tsai, Jing-fu}
}
@article {1507,
	title = {Historical biogeography of scarabaeine dung beetles},
	journal = {Journal of Biogeography},
	volume = {29},
	number = {9},
	year = {2002},
	note = {598JTJ BIOGEOGR},
	month = {Sep},
	pages = {1217-1256},
	abstract = {Aim (1) To review briefly global biogeographical patterns in dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae), a group whose evolutionary history has been dominated by ecological specialization to vertebrate dung in warmer climates. (2) To develop hypotheses accounting for the evolution of these patterns. Location Six principal biogeographical regions: Palaearctic, Oriental, Afrotropical, Australasia, Neotropical, Nearctic and five outlying islands or island groups harbouring endemic genera: Caribbean, Madagascar, Mauritius, New Caledonia, New Zealand. Methods Major patterns of tribal, generic and species distribution are investigated using cluster analysis, ordination, parsimony analysis of endemism and track analysis. Attempts are made to resolve biogeographical patterns with findings in the fields of plate tectonics, fossil and evolutionary history, plus phylogeny of both mammals and dung beetles. Results Because of conflict between published findings, it is uncertain at what point in time density of dinosaur dung, mammal dung or both became sufficiently great to select for specialized habits in dung beetles. However, biogeographical evidence would suggest a Mesozoic origin followed by further taxonomic radiation during the Cenozoic, possibly in response to the increasing size and diversity of mammalian dung types in South America and Afro-Eurasia. Proportional generic distribution in fourteen tribes and subtribes showed four principal biogeographical patterns: (1) southerly biased Gondwanaland distribution, (2) Americas or (3) Madagascar endemism, and (4) northerly biased, Afro-Eurasian- centred distribution with limited numbers of genera also widespread in other regions. Proportional composition of faunas in eleven geographical regions indicated three principal distributional centres, East Gondwanaland fragments, Afro- Eurasia and the Americas. These patterns probably result from three principal long-term range expansion and vicariance events (Mesozoic: Gondwanaland interchange and fragmentation, Cenozoic: Afro-Eurasian/Nearctic interchange and the Great American interchange). It is suggested that old vicariance caused by the Mesozoic fragmentation of Gondwanaland leads to a high degree of regional endemism at generic or tribal level across one or more Gondwanaland tracks. In contrast, it is suggested that the more recent Cenozoic range expansions occurred primarily towards northern regions leading to endemism primarily at species level. These Cenozoic radiations were facilitated by the re-linking of continents, either because of tectonic plate movements (Africa to Eurasia in Miocene), climatically induced sea-level change (Afro-Eurasia to Nearctic in Miocene and Pleistocene), or similar coupled with orogenics (Nearctic to Neotropical in Pliocene). Speciation has followed vicariance either because of climatic change or physical barrier development. These recent range expansions probably occurred principally along an Afro-Eurasian land track to the Nearctic and Neotropical and an Americas land track northwards from the Neotropics to the Nearctic, with limited dispersal from Eurasia to Australia, probably across a sea barrier. This accounts for the overall, spatially constrained, biogeographical pattern comprising large numbers of species- poor genera endemic to a single biogeographical region and fewer more species-rich genera, many of which show wider biogeographical distributions. In most southerly regions (Australasia, Madagascar, Neotropical), faunal composition and generic endemism is primarily dominated by elements with Gondwanaland ancestry, which is consistent with the Gondwanaland origin claimed for Scarabaeinae. In Afro-Eurasia (Palaearctic, Oriental, Afrotropical), generic endemism of monophyletically derived Afro-Eurasian and widespread lineages is centred in the Afrotropical region and faunal composition is numerically dominated by Afro-Eurasian and widespread elements. In the Nearctic region, the fauna is jointly dominated by widespread elements, derived from Afro-Eurasia, and Gondwanaland and Americas elements derived from the Neotropical region. Main conclusions Global biogeographical patterns in scarabaeine dung beetles primarily result from Mesozoic and Cenozoic range expansion events followed by vicariance, although recent dispersal to Australia may have occurred across sea barriers. Detailed phylogenetics research is required to provide data to support dispersal/vicariance hypotheses.},
	keywords = {assemblage coleoptera, biogeography, community organization, dinosaurs, dispersal, distributional patterns, Dung, fragmentation, global, habitat, hindwing articulation, historical, mammals, rain-forest, s-str, Scarabaeinae, southern african, vicariance, w},
	url = {<Go to ISI>://000178273300010},
	author = {Davis, A. L. V. and Scholtz, C. H. and Philips, T. K.}
}
@article {1715,
	title = {Spatial distribution of three groups of Coleoptera along an altitudinal transect in the Mexican Transition Zone and its biogeographical implications},
	journal = {Elytron},
	volume = {9},
	year = {1995},
	pages = {151-185},
	keywords = {Altitudinal distribution, biogeography, COLEOPTERA, Geotrupidae, Scarabaeinae, Silphidae, Veracruz},
	isbn = {0214-1353},
	author = {Halffter, Gonzalo and Favila, Mario E and Arellano, Lucrecia}
}
@article {1790,
	title = {Dung beetle geographic diversity variation along a Western Iberian latitudinal transect (Coleoptera: Scarabaeidae)},
	journal = {Annals of the Entomological Society of America},
	volume = {93},
	number = {2},
	year = {2000},
	pages = {235-243},
	abstract = {To explore the richness and geographic turnover of dung beetle assemblages (Scar- abaeinae and Aphodiinae) inhabiting the Western Iberian peninsula, a latitudinal north{\DH}south transect was carried out in Portuguese territory. Estimators of alpha, beta, and gamma diversities were used, as well as the ratio Scarabaeinae/Aphodiinae, which puts into evidence some dung beetle community spatial characteristics. Clustering methods were also used to ascertain relationships between biogeographical Portuguese regions and geographical patterns of species replacement. Although Scarabaeinae appears mainly responsible for overall and regional richness, Aphodiinae richness was rather low everywhere except for the southernmost plot of the transect investigated. The scarcity of Aphodiinae species is re{\ss}ected in the Scarabaeinae/Aphodiinae ratio, a score that is, however, clearly biased toward the Aphodiinae in almost all the Iberian Peninsula. For this reason the pairwise turnover in species composition between sites is mainly determined by the variation in Scarabaeinae species. Peaks of },
	keywords = {Aphodiinae, biogeography, Dung beetles, faunistic turnover, Portugal, Scarabaeinae},
	author = {Hortal-Munoz, J. and Martin-Pierra, Fermin and Lobo, Jorge M.}
}
@article {2052,
	title = {Biogeographic areas and transition zones of Latin America and the Caribbean islands based on panbiogeographic and cladistic analyses of the entomofauna},
	journal = {Annual Review of Entomology},
	volume = {51},
	year = {2006},
	pages = {467-494},
	abstract = {Track and cladistic biogeographic analyses based on insect taxa are used as a framework to interpret patterns of the Latin American and Caribbean ento- mofauna by identifying biogeographic areas on the basis of endemicity and arranging them hierarchically in a system of regions, subregions, dominions, and provinces. The Nearctic region, inhabited by Holarctic insect taxa, comprises five provinces: California, Baja California, Sonora, Mexican Plateau, and Tamaulipas. The Mexi- can transition zone comprises five provinces: Sierra Madre Occidental, Sierra Madre Oriental, Transmexican Volcanic Belt, Balsas Basin, and Sierra Madre del Sur. The Neotropical region, which harbors many insect taxa with close relatives in the tropical areas of the Old World, comprises four subregions: Caribbean, Amazonian, Chacoan, and Parana. The South American transition zone comprises five provinces: North An- dean Paramo, Coastal Peruvian Desert, Puna, Atacama, Prepuna, and Monte. The An- dean region, which harbors insect taxa with close relatives in the Austral continents, comprises three subregions: Central Chilean, Subantarctic, and Patagonian. . R },
	keywords = {biogeography, biotic components, evolution, insect endemicity, provinces, regions},
	author = {Morrone, Juan J.}
}
@article {2053,
	title = {Halffter{\textquoteright}s mexican transition zone, beetle generalized tracks, and geographical homology},
	journal = {Journal of Biogeography},
	volume = {28},
	year = {2001},
	pages = {635-650},
	keywords = {biogeography, COLEOPTERA, homology, mesoamerica, Neotropics, panbiogeography},
	author = {Morrone, Juan J. and Marquez, Juan}
}
@article {2213,
	title = {Phylogeography of the Namib Desert dung beetles Scarabaeus (Pachysoma) MacLeay (Coleoptera: Scarabaeidae)},
	journal = {Journal of Biogeography},
	volume = {32},
	year = {2005},
	pages = {75-84},
	abstract = {Aim Namib biogeography in many instances remains reliant on advanced and detailed systematic studies. This study attempts to combine molecular phylogenetic data, geology and palaeo-climatic data to (i) resolve the relationships of the 13 morphological species of Scarabaeus (Pachysoma) and (ii) relate their evolution to past climatic and geological events. Location South Africa and Namibia. Methods Sequencing of a 1197 bp segment of the mitochondrial cytochrome oxidase I (COI) gene of the 13 species within Scarabaeus (Pachysoma) was undertaken. Analyses performed included Parsimony and Maximum Likelihood as well as imposing a molecular clock. Results The molecular phylogeny showed strong support for 11 of the 13 morphological species. The remaining two species, S. (P.) glentoni and S. (P.) hippocrates, formed a complex and could not be assigned specific status on the basis of the COI gene phylogeny. Strong support for the three species formerly classified within the genus Neopachysoma was consistently obtained. The subgenus appears to have arisen c. 2.9 Ma. Species within the subgenus arose at different times, with the common ancestor to Neopachysoma and the hippocrates complex having evolved 2.65 and 2.4 Ma, respectively. Scarabaeus (P.) denticollis, S. (P.) rotundigenus, S. (P.) rodriguesi and S. (P.) schinzi are some of the youngest species, having diverged between 2 million and 600,000 years ago. Main conclusions Scarabaeus (Pachysoma) is a derived monophyletic clade within the Scarabaeini. The subgenus appears to be young in comparison with the age of the Namib Desert, which dates back to the Miocene (c. 15 Ma). The psammophilous taxa are shown to disperse with their substratum and habitat, barchan dunes. Clear south/north evolutionary gradients can be seen within the species of this subgenus, which are consistent with the unidirectional wind regime. Species with a suite of mostly plesiomorphic characters have a southerly distribution while their derived psammophilous relatives have central to northern Namib distributions. Major rivers such as the Orange, Buffels and Holgat appear to be gene barriers to certain species as well as areas of origin of speciation events. },
	keywords = {Aptery, biogeography, COLEOPTERA, cytochrome oxidase I, endemic, mitochondrial DNA, namib desert, phylogeny, Scarabaeidae, Scarabaeus (Pachysoma).},
	author = {Sole, Catherine L. and Scholtz, Clarke H. and Bastos, Armanda D. S.}
}
