@article {1332,
	title = {Diversity of dung and carrion beetles in a disturbed Mexican tropical montane cloud forest and on shade coffee plantations},
	journal = {Biodiversity and Conservation},
	volume = {14},
	number = {3},
	year = {2005},
	note = {digital copy},
	pages = {601-615},
	abstract = {This paper analyzes the diversity of dung and carrion beetles (Scarabaeinae and Silphidae) in four human-induced habitats of a disturbed tropical montane cloud forest: polyspecific shade coffee plantations, monospecific shade coffee plantations, tropical montane cloud forest fragments, and clear cuts. The four habitats had similar richness, species composition, and assemblage structure of dung and carrion beetles. Differences were found in abundance and biomass levels for the four dominant species in the landscape. Dung beetles were more abundant than carrion beetles, but the biomass was higher for the latter. Carrion beetles were seasonal, while dung beetles were clearly not. When forest fragments and shade coffee plantations were compared to other similar habitats in the region, the same general pattern was observed. However, forests with high disturbance and monospecific shade coffee plantations had lower species richness than forests with low and medium disturbance and polyspecific shade coffee plantations. Thus shade coffee plantations maintain connectivity between patches of cloud forest in a landscape that is strongly affected by human activities. Protecting landscape diversity appears to ensure high species richness.},
	keywords = {anthropogenic change, biodiversity, carrion beetles, dung beetle, fragmentation, modification (biological conservation 2007), shade coffee plantations, tropical montane cloud forest, Veracruz},
	author = {Arellano, Lucrecia and Favila, Mario E and Huerta, Carmen}
}
@article {1333,
	title = {Gamma diversity: derived from and a determinant of alpha diversity and beta diversity. An analysis of three tropical landscapes},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {90},
	year = {2003},
	pages = {27-76},
	abstract = {Using three taxonomic groups of beetles we examine how alpha and beta diversity influence the species richness of a landscape (gamma diversity), and vice versa. That is, how the species richness of a landscape {\textendash} which is a historical and biogeographical phenomenon {\textendash} contributes to the values of alpha diversity (1) at a given site, (2) in a community, (3) in terms of cumulative species richness by community, and also contributes to (4) the intensity of species exchange between communities. To explore this question, we used two subfamilies of Scarabaeoidea: Scarabaeinae and Geotrupinae, and the family Silphidae. In all analyses these three taxonomic groups are considered as a single indicator group: the copronecrophagous beetles. Information is also included on the subfamily Aphodiinae (Scarabaeoidea), coprophagous beetles not included in the indicator group. Several types of vegetation located in three landscapes (tropical, transition and mountain) were studied, and these are located along an altitudinal gradient in the central part of the state of Veracruz, Mexico. We base this study on the following concepts. The alpha diversity of an indicator group reflects the number of species that use a given environment or resource in a given place or community. Spacial beta diversity is related to the response of organisms to spatial heterogeneity. Gamma diversity depends primarily on the historical and geographic processes that operate on the mesoscale level and is also affected by alpha and beta diversity. It is on this scale of landscape that human actions, such as the modification and fragmentation of vegetation, have their most important effects. These are, however, often beyond the scope of ecological analyses carried out on a local scale. In the three landscapes, sampling was carried out regularly at 67 sites, with complementary sampling at another 69 sites. Twenty-six types of vegetation communities were studied. A total of 16,152 specimens representing 60 species were captured (52 species of Scarabaeinae, 4 Geotrupinae and 4 Silphidae). In the tropical landscape the community richest in species was low deciduous forest. In the transition landscape, cloud forest was the richest. Each of these communities is the most representative of their respective altitudinal bands. In contrast, the greatest species richness in the mountain landscape occurred in the mountain grasslands and pastures; types of community favoured by or even created by human intervention. This is explained by the expansion of heliophilous species from the Mexican High Plateau into these areas. In the tropical landscape the species richness of the pastures is similar to that of its forests, but with a partially different composition which is characterized by the dominance of heliophilous and coprophagous species; the latter, in addition to the more ubiquitous species that are shared with the tropical forest. In the transition landscape the cloud forest and the coffee plantations with polyspecific shade are important in the context of conserving the fauna. This type of community offers arboreal cover and occupies the majority of this landscape, allowing the groups of insects studied to move between remnant fragments of cloud forest. On the landscape scale but not locally, the fragmentation of natural communities does not appear to have reduced the number of species for the beetles of the indicator group. In each landscape disturbance by human activity appears to have been overcome for distinct reasons. In the tropical landscape we find the heliophilous beetle fauna characteristic of pastures, and this has increased by two species of recent invaders. In the transition landscape, the coffee plantations with polyspecific shade create a communication matrix, while in the mountain landscape the expansion of the mountain pastures has made conditions more favourable for heliophilous species. These results are not necessarily expected for other groups of organisms. },
	keywords = {alpha, anthropogenic landscape, beta and gamma diversity, Dung beetles, Geotrupinae, Scarabaeinae, Silphidae, Veracruz},
	author = {Arellano, Lucrecia and Halffter, Gonzalo}
}
@article {1334,
	title = {Response of dung beetle assemblages to landscape structure in remnant natural and modified habitats in Southern Mexico},
	journal = {Insect Conservation and Diversity},
	volume = {1},
	year = {2008},
	pages = {253-262},
	abstract = {1. To provide reliable tools for landscape management in tropical regions, it is important to recognise the emergent properties of landscape heterogeneity and to understand their influence on different components of biodiversity. 2. The effects of habitat type, patch size and shape, canopy cover, distance between habitats, and elevation were examined on the partitioned diversity (alpha, beta and gamma) of dung beetles in a transformed landscape in southern Mexico. 3. In total, 4109 individuals belonging to 28 species of dung beetle were recorded in the entire landscape. Alpha diversity was similar among habitats, although species com- position was more similar between forest remnants and living fences than between living fences and pastures. Beta diversity was related to differences in elevation and distance between habitats. The distance between fragments was significantly correlated with the mean number of beetle captures and species richness per site. Gamma diversity depended more on species turnover (beta diversity) than on local species richness. 4. In general, living fences and pastures with trees, and small forest patches retain a significant proportion of the fauna typically associated with pristine forest habitats. The remaining tracts of forest, along with the contemporary landscape elements may offer suitable habitats for future recolonisation by other insect species in heavily disturbed landscapes in tropical Mexico.},
	keywords = {Alpha and beta diversity, biodiversity, indicator groups, landscape management, living fences, pastures, Scarabaeinae, tropical dry forest.},
	author = {Arellano, Lucrecia and Leon-Cortes, Jorge L. and Halffter, Gonzalo}
}
@article {1335,
	title = {Patterns of abundance and movement in relation to landscape structure: a study of a common scarab (Canthon cyanellus cyanellus) in Southern Mexico},
	journal = {Landscape Ecology},
	volume = {23},
	year = {2008},
	pages = {69-78},
	keywords = {agro-pasture systems, animal movement, Chiapas, diffusion, functional connectivity, hedgerow, Scarabs, state-space model, tropical deciduous forest},
	author = {Arellano, Lucrecia and Leon-Cortes, Jorge L. and Ovaskainen, Otso}
}
@article {1586,
	title = {From forest to pasture: an evaluation of the influence of environment and biogeography on the structure of dung beetle (Scarabaeinae) assemblages along three altitudinal grafients in the Neotropical region},
	journal = {Ecography},
	volume = {30},
	year = {2007},
	month = {Feb},
	pages = {193-208},
	abstract = {The objective of this study is to evaluate the effect of environmental (associated with the expansion of cattle ranching) and biogeographical factors on the diversity of dung beetle (Scarabaeinae) assemblages along three altitudinal gradients in the Neotropical region. One gradient is located in the Mexican Transition Zone, on the Cofre de Perote mountain, the other two are in the northern Andes (the Chiles Volcano and the R{\i} ́o Cusiana Basin). For the three gradients, the number of species decreased as altitude increased. On the Cofre de Perote, regardless of altitude, the number of species and of individuals was similar in both forest and pasture, while species composition was different between habitats. On this mountain, species turnover in pastures was characterized by the addition of new species as altitude increased. In the northern Andes, species diversity was always greater in the forest than in the pasture, and species turnover between habitats was notably influenced by species loss with increasing altitude. As such the pasture fauna of the northern Andes was an impoverished derivative of the fauna present in the forests at the same altitude characterized by species of Neotropical affinity with a limited capacity for colonizing open, sunnier habitats. The opposite occurs in the areas used by cattle on the Cofre de Perote. This habitat has its own fauna, which is mainly comprised of Holarctic and Afrotropical species adapted to the prevailing environmental conditions of areas lacking arboreal vegetation. These results suggest that the impact on beetle communities caused by human activities can differ depending on the geographic position of each mountain and, particularly, the biogeographical history of the species assemblage that lives there.},
	author = {Escobar S. , Federico and Halffter, Gonzalo and Arellano, Lucrecia}
}
@inbook {1711,
	title = {Variaci{\'o}n de la diversidad en especies de Scarabaeinae (Coleoptera:Scarabaeidae) como respuesta a la antropizaci{\'o}n de un paisaje tropical},
	booktitle = {T{\'o}picos sobre Coleoptera de M{\'e}xico},
	year = {2001},
	note = {have a copy},
	pages = {35-53},
	publisher = {Universidad de Guadalajara{\textemdash}Universidad Aut{\'o}noma del Estado de Morelos},
	organization = {Universidad de Guadalajara{\textemdash}Universidad Aut{\'o}noma del Estado de Morelos},
	address = {Guadelajara, M{\'e}xico},
	keywords = {community, conservation (biological conservation 2007), deforestation, diversity, microclimate, modification, Scarabaeinae},
	author = {Halffter, Gonzalo and Arellano, Lucrecia},
	editor = {Navarrete-Heredia, J. L. and Fierros Lop{\'e}z, H. E. and Burgos-Solorio, A.}
}
@article {1712,
	title = {Response of dung beetle diversity to human-induced changes in a tropical landscape},
	journal = {Biotropica},
	volume = {34},
	number = {1},
	year = {2002},
	note = {have a copydigital copy too},
	pages = {144-154},
	keywords = {disturbance mammals, indicator, Mexico, Scarabaeinae, tropical decicuous forest},
	author = {Halffter, Gonzalo and Arellano, Lucrecia}
}
@article {1715,
	title = {Spatial distribution of three groups of Coleoptera along an altitudinal transect in the Mexican Transition Zone and its biogeographical implications},
	journal = {Elytron},
	volume = {9},
	year = {1995},
	pages = {151-185},
	keywords = {Altitudinal distribution, biogeography, COLEOPTERA, Geotrupidae, Scarabaeinae, Silphidae, Veracruz},
	isbn = {0214-1353},
	author = {Halffter, Gonzalo and Favila, Mario E and Arellano, Lucrecia}
}
@article {1721,
	title = {Instability of copronecrophagous beetle assemblages (Coleoptera: Scarabaeinae) in a mountainous tropical landscape of Mexico},
	journal = {Environmental Entomology},
	volume = {36},
	number = {6},
	year = {2007},
	pages = {1397-1407},
	abstract = {We analyzed changes over time in species composition and functional guild structure (temporal beta diversity) for natural assemblages and those modi{\TH}ed by humans in a fragmented, tropical mountain landscape. The assemblages belong to cloud forests (the original vegetation type), secondary forests, traditional shaded coffee plantations, commercial shaded coffee plantations, and a cattle pasture. Copronecrophagous beetles, subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae), were used as the indicator group. This group has been used in previous studies and other tropical forests and has been found to be a good indicator of the effects of anthropogenic change. For each assemblage, we compared samples that were collected several years apart. Changes were found in species composition, order of abundance, and in the proportion that a given species is present in the different functional groups. The changes that occurred between samplings affected the less abundant species in the cloud forest and in the pasture. In the other vegetation types, both abundant and less abundant species were affected. Their order of abundance and proportion in the different guilds also changed. This study shows that, although landscape richness remains relatively constant, richness at the local level (alpha diversity) changes notably even over short lapses of time. This could be a characteristic of landscapes with intermediate degrees of disturbance (such as those that have been partially modi{\TH}ed for human use), where assemblage composition is very fluid. },
	keywords = {alpha diversity, beta diversity, cloud forest, fragmented landscapes, Scarabaeinae},
	author = {Halffter, Gonzalo and Pineda, Eduardo and Arellano, Lucrecia and Escobar S. , Federico}
}
@article {2274,
	title = {Thermoregulation in endothermic dung beetles (Coleoptera: Scarabaeidae): effect of body size and ecophysiological constraints in flight},
	journal = {Journal of Insect Physiology},
	volume = {52},
	year = {2006},
	pages = {854-860},
	abstract = {We explore the physiological constraints of body temperature as related to body mass and ambient temperature during flight in endothermic dung beetles showing a mass-related breakpoint where species show strong vs. weak endothermy. We found two different strategies in the dung beetles prior to flight; larger beetles (41.9 g) elevate and maintain their body temperature (Tb) at levels well above ambient temperature (Ta) whereas smaller beetles{\textquoteright} (o1.9 g) Tb tends to conform with Ta. Physiological constraints analysis revealed a constant maximum tolerated temperature (in flight) of 42 1C and a minimum temperature for flight of around 25 1C. These, with body mass, may play a role in thermal niche partitioning and geographical distribution patterns. },
	keywords = {body size, Ecophysiology, Null models, Scarabaeoidea., Scarabs, thermoregulation},
	author = {Verdu, J. R. and Arellano, Lucrecia and Numa, C.}
}
@article {2275,
	title = {Roles of endothermy in niche differentiation for ball-rolling dung beetles (Coleoptera: Scarabaeidae) along an altitudinal gradient},
	journal = {Ecological Entomology},
	volume = {32},
	year = {2007},
	pages = {544-551},
	abstract = {    1.   An analysis of whether niche differentiation in ball-rolling dung beetles can be explained by the way in which they regulate their body temperature was conducted.  2.    A priori  assumptions were: (i) if thermoregulation affects niche partitioning, sympatric species must have different endothermic strategies that minimise encounters; or, alternatively (ii) if two co-occurring species show the same thermoregulation pattern and their flight periods overlap, they might be avoiding competition by exhibiting different resource preferences or different food relocation behaviour.  3.   The ball-rolling dung beetles studied showed a hierarchical structure based on the species{\textquoteright} endothermic capacity, measured as temperature excess [ T  ex  = difference between body temperature ( T  b ) and ambient temperature ( T  a )]. Those with a high  T  ex  (10 {\textendash} 15 {\textdegree}C) were located exclusively at altitudes >1000 m a.s.l. On the coastal plains, species with a high  T  ex  were restricted to flying at night when the  T  a  was lower. Species with a lower  T  ex  (less than 10 {\textdegree}C higher than  T  a ) were found in the coastal plains zone.  4.   Where there was sympatry with similar trophic habits, the species involved showed very different thermal niches, and where there was significant overlap of thermal niches between sympatric species, trophic habits of species were very different.  5.   The results suggest that it is possible to use the concept of the thermal niche as a tool to explain interspecific interactions and the spatial distribution of species.  },
	keywords = {Ecophysiology, niche overlap, Null models, Scarabaeoidea  ., thermal niche, thermoregulation},
	author = {Verdu, Jose R. and Arellano, Lucrecia and Numa, Catherine and Mico, Estefania}
}
