@article {1309,
	title = {Comparaci{\'o}n de la composici{\'o}n y riqueza de especies de escarabajos copr{\'o}fagos (Coleoptera: Scarabaeidae) en remanentes de bosque de la Orinoquia Colombiana},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {76},
	year = {1999},
	note = {have copy},
	pages = {113-126},
	keywords = {Colombia, community, deforestation, dung beetle (biological conservation 2007), fragmentation, richness, Scarabaeinae},
	author = {Amezquita, Sandra J. and Forsyth, Adrian and Lopera, Alejandro and Camacho, Andr{\'e}s}
}
@article {1325,
	title = {Escarabajos Coprofagos (Coleoptera: Scarabaeoidea) asociados a hongos en la Sierra Madre Occidental, Durango, Mexico: con una compilacion de las especies micetofagas},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {80},
	year = {2000},
	pages = {119-130},
	keywords = {Durango, Mexico, Mycetophagy, Scarabaeidae},
	author = {Sofia Anduaga}
}
@article {1327,
	title = {Nuevos registros de escarabajos coprofagos (Coleoptera: Scarabaeidae) en detritus de madrigueras de Neotoma albigula Hartley (Rodentia: Muridae)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {23},
	number = {3},
	year = {2007},
	pages = {143-144},
	abstract = {The presence of two species of dung beetles (Scarabaeidae: Scarabaeinae): Onthophagus browni Howden and Cartwright and Copris macclevei Warner are mentioned for the first time in nests of vertebrates: Neotoma albigula Hartley in Durango, Mexico. },
	author = {Sofia Anduaga}
}
@article {1330,
	title = {Nidificacion y alimentacion en Liatongus rhinocerulus (Bates) (Coleoptera: Scarabaeidae: Scarabaeinae)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {57},
	year = {1993},
	pages = {1-14},
	keywords = {behavior, Liatongus, Mycetophagy, nidification, Scarabaeinae},
	author = {Sofia Anduaga and Halffter, Gonzalo}
}
@article {1333,
	title = {Gamma diversity: derived from and a determinant of alpha diversity and beta diversity. An analysis of three tropical landscapes},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {90},
	year = {2003},
	pages = {27-76},
	abstract = {Using three taxonomic groups of beetles we examine how alpha and beta diversity influence the species richness of a landscape (gamma diversity), and vice versa. That is, how the species richness of a landscape {\textendash} which is a historical and biogeographical phenomenon {\textendash} contributes to the values of alpha diversity (1) at a given site, (2) in a community, (3) in terms of cumulative species richness by community, and also contributes to (4) the intensity of species exchange between communities. To explore this question, we used two subfamilies of Scarabaeoidea: Scarabaeinae and Geotrupinae, and the family Silphidae. In all analyses these three taxonomic groups are considered as a single indicator group: the copronecrophagous beetles. Information is also included on the subfamily Aphodiinae (Scarabaeoidea), coprophagous beetles not included in the indicator group. Several types of vegetation located in three landscapes (tropical, transition and mountain) were studied, and these are located along an altitudinal gradient in the central part of the state of Veracruz, Mexico. We base this study on the following concepts. The alpha diversity of an indicator group reflects the number of species that use a given environment or resource in a given place or community. Spacial beta diversity is related to the response of organisms to spatial heterogeneity. Gamma diversity depends primarily on the historical and geographic processes that operate on the mesoscale level and is also affected by alpha and beta diversity. It is on this scale of landscape that human actions, such as the modification and fragmentation of vegetation, have their most important effects. These are, however, often beyond the scope of ecological analyses carried out on a local scale. In the three landscapes, sampling was carried out regularly at 67 sites, with complementary sampling at another 69 sites. Twenty-six types of vegetation communities were studied. A total of 16,152 specimens representing 60 species were captured (52 species of Scarabaeinae, 4 Geotrupinae and 4 Silphidae). In the tropical landscape the community richest in species was low deciduous forest. In the transition landscape, cloud forest was the richest. Each of these communities is the most representative of their respective altitudinal bands. In contrast, the greatest species richness in the mountain landscape occurred in the mountain grasslands and pastures; types of community favoured by or even created by human intervention. This is explained by the expansion of heliophilous species from the Mexican High Plateau into these areas. In the tropical landscape the species richness of the pastures is similar to that of its forests, but with a partially different composition which is characterized by the dominance of heliophilous and coprophagous species; the latter, in addition to the more ubiquitous species that are shared with the tropical forest. In the transition landscape the cloud forest and the coffee plantations with polyspecific shade are important in the context of conserving the fauna. This type of community offers arboreal cover and occupies the majority of this landscape, allowing the groups of insects studied to move between remnant fragments of cloud forest. On the landscape scale but not locally, the fragmentation of natural communities does not appear to have reduced the number of species for the beetles of the indicator group. In each landscape disturbance by human activity appears to have been overcome for distinct reasons. In the tropical landscape we find the heliophilous beetle fauna characteristic of pastures, and this has increased by two species of recent invaders. In the transition landscape, the coffee plantations with polyspecific shade create a communication matrix, while in the mountain landscape the expansion of the mountain pastures has made conditions more favourable for heliophilous species. These results are not necessarily expected for other groups of organisms. },
	keywords = {alpha, anthropogenic landscape, beta and gamma diversity, Dung beetles, Geotrupinae, Scarabaeinae, Silphidae, Veracruz},
	author = {Arellano, Lucrecia and Halffter, Gonzalo}
}
@article {1432,
	title = {Fauna de Coleoptera Scarabaeoidea de Cuetzalan del Progreso, Puebla, Mexico},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {88},
	year = {2003},
	pages = {87-121},
	abstract = {In this work a comparative analysis of the scarab beetles of Cuetzalan del Progreso, Puebla, México is presented. Data were obtained from March 1998 to March 1999 in tropical rain forest, induced pasture land and secondary vegetation, between 400-1000 m above sea level. Records of 7 subfamilies, 18 subtribus, 34 genera and 63 species are presented. The greatest specific diversity corresponds to Dynastinae (21 spp), Melolonthinae (13 spp), Scarabaeidae (13 spp) and Passalinae (8 spp). The 50\% of the species are concentrated in the genera: Phyllophaga (9 spp), Cyclocephala (9 spp), Onthophagus (5 spp), Ataenius (3 spp), Passalus (3 spp) and Anomala (2 spp). We present a parsimony analysis of endemicity in order to know the relationships between the fauna of Cuetzalan and other Mexican localities. The fauna of Cuetzalan de Progreso is closely related to the fauna of the humid mountains of the northeast of Hidalgo. A key to the taxa found in Cuetzalan del Progreso is included. },
	keywords = {diversity, Mexico, Scarabaeidae, Sierra Madre Oriental, taxonomy},
	author = {Carrillo-Ruiz, Hortensia and Moron, Miguel-Angel}
}
@article {1440,
	title = {Primer registro de Canthon angustatus Harold 1867 en Mexico (Coleoptera: Scarabaeoidea)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {21},
	number = {3},
	year = {2005},
	pages = {159-160},
	abstract = {The scarab beetle Canthon angustatus Harold, 1867 (Coleoptera: Scarabaeoidea) is recorded for the first time in Mexico, based upon several specimens collected in Chiapas. },
	author = {Chame-Vazquez, Eduardo R. and Gomez y Gomez, Benigno}
}
@article {1514,
	title = {Estado del conocimiento y conservacion de los coleopteros Scarabaeoidea (Insecta) del estado de Hidalgo, Mexico},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {22},
	number = {2},
	year = {2006},
	pages = {57-108},
	abstract = {A faunistic and conservation-oriented analysis of the Scarabaeoidea beetles from the Hidalgo State, México is presented. Seven families with 95 genera and 276 species are recorded, of which five genera and six species are considered introduced. Fifteen species are cited for the first time for this state and 21 species are considered to be erroneously or doubtfully recorded for Hidalgo. Thus, Hidalgo is the fifth Mexican state with the highest number of Scarabaeoidea species. On the basis of estimates of species numbers of several biological groups, it is extrapolated that at least 100 species are missing to record from this state. According to the geographical distribution of the species, six groups are proposed (with the exception of introduced species): endemic from Hidalgo (13 spp.), endemic from México (104 spp.), North American (37 spp.), Central American (55 spp.), Latin American (32 spp.), and American (29 spp.). With respect to the vegetation types of the state, the coniferous forest has 92 species of which 37 are restricted to this forest, the oak forest 71 species and none are restricted, the cloud forest 108 species with 34 restricted, the tropical evergreen and semideciduous forests 108 species with 29 restricted, the tropical deciduous forest 69 species with one restricted, and the xerophilous scrub 52 species with 12 restricted. The distribution of the species by biogeographical provinces shows that 62.1 \% of the total species are exclusively located in the Sierra Madre Oriental, 13.8 \% in the Altiplano Mexicano, 3.8 \% in the Eje Neovolcánico Transmexicano and 1.9 \% in the Golfo de México. With regard to endemism of Hidalgo and México, the coniferous forest presents 60 species, the cloud forest 59 species, the oak forest 44 species, the tropical evergreen and semideciduous forests 16 species, the xerophilous scrub 13 species, and the tropical deciduous forest nine species. With regard to the four federal protected natural areas of the Hidalgo state, it is necessary to carry out a systematic faunal work at the Biosphere Reserve Barranca de Metztitlán, due to the lack of data of scarab beetles from this and many other arid and semiarid areas of México. It is also important to complete the faunal inventories of the areas with coniferous and oak forests, among them the National Parks Los Mármoles and El Chico. In addition, it is advisable to conduct supplementary collections in areas with tropical deciduous forest, since information on scarab beetles in this forest in Hidalgo state is scarce. On the basis of the high values of richness, endemism and restricted species numbers of the scarab beetles inhabiting the cloud forests, it is urgent to propose protected areas in the state of Hidalgo with this type of vegetation. It is also necessary to obtain more information on the eco-geographical distribution of scarab beetles from Hidalgo, in order to propose species into categories of risk. Likewise, the present-day number of protected natural areas is considered inadequate to preserve this insects group.},
	keywords = {COLEOPTERA, conservation, endemism, Hidalgo, Me, protected natural areas, Scarabaeoidea, taxonomic diversity, vegetation types, ́xico},
	author = {Delgado, Leonardo and Marquez, Juan}
}
@article {1516,
	title = {Necrophilous Scarabaeidae and Trogidae beetles of tropical deciduous forest in Tepexco, Puebla, Mexico},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {52},
	year = {1992},
	pages = {1-13},
	keywords = {COLEOPTERA, Mexico, necrophagous, Scarabaeidae, Trogidae, tropical deciduous forest},
	author = {DeLoya, Cuauhtemoc}
}
@article {1581,
	title = {Diversidad de Coleopteros coprofagos (Scarabaeidae: Scarabaeinae) en un mosaico de habitats en la reserva natural Nunak, Guaviare, Colombia},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {79},
	year = {2000},
	pages = {103-121},
	keywords = {Amazon, Colombia, diversity, dung beetle, Scarabaeinae, spatial heterogeneity},
	author = {Escobar S. , Federico}
}
@article {1582,
	title = {Seleccion de habitat y comportamiento sexual de Sulcophanaeus velutinus (Murray 1856) (Coleoptera: Scarabaeidae: Scarabaeinae) en un bosque de montana en Colombia},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {90},
	year = {2003},
	pages = {307-310},
	author = {Escobar S. , Federico}
}
@article {1697,
	title = {Estados preimaginales, nidificacion y fenologia de Canthidium (E.) moestum Harold, 1867 (Coleoptera, Scarabaeidae, Coprini)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {73},
	year = {1998},
	pages = {155-165},
	keywords = {canthidium moestum, COLEOPTERA, Coprinae, larva, life cycle, nesting behavior, phenology, pupa, Scarabaeidae, taxonomy},
	author = {Gonzalez-Vainer, Patricia and Morelli, Enrique}
}
@article {1755,
	title = {Desarrollo larval de Canthon cyanellus cyanellus LeConte 1859 (Coleoptera: Scarabaeidae)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {89},
	year = {2003},
	pages = {185-200},
	abstract = {The larval development of Canthon cyanellus cyanellus LeConte was analyzed upon 92 larvae of known age. Larval stage lasted nine days and consists of three instars; each one with a particular length of time and specific cephalic-width range. Most of the anatomical larval features appear since the first instar and they strengthen in the following two. The comparison of several anatomical features among larvae of Canthon cyanellus cyanellus, C. pilularius Linnaeus, C. humectus humectus Say and C. edentulus Harold showed that C. cyanellus cyanellus and C. pilularius, shared the most numerous characters in common wich could indicate that these species are taxonomically close. },
	keywords = {anatomy, Canthon cyanellus, larval instars, morphometry, Scarabaeidae},
	author = {Hernandez-Martinez, Gerardo and Martinez M. , Imelda}
}
@article {1811,
	title = {Comparative analysis of reproductive and nesting behavior in several species of Eurysternus Dalman (Coleoptera: Scarabaeinae: Eurysternini)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {88},
	year = {2003},
	pages = {1-41},
	keywords = {Dung beetles},
	author = {Huerta, Carmen and Halffter, Gonzalo and Violeta Halffter and L{\'o}pez, Rosario}
}
@article {1862,
	title = {A preliminary study of the invasion and dispersal of Digitonthophagus gazella (Fabricius, 1787) in Mexico (Coleoptera: Scarabaeidae: Scarabaeinae)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {61},
	year = {1994},
	pages = {35-42},
	keywords = {dispersal, introduced fauna, Mexico, Scarabaeinae},
	author = {Kohlmann, Bert}
}
@article {1927,
	title = {Relaciones entre escarabajos (Coleoptera: Scarabaeidae) y nidos de tuza (Rodentia: Geomydae): implicaciones biologicas y biogeograficas},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {62},
	year = {1994},
	pages = {1-9},
	keywords = {{\textquoteright}, association with rodents, nests, Onthophagus, Scarabaeinae},
	author = {Lobo, Jorge M. and Halffter, Gonzalo}
}
@article {1937,
	title = {Spatial microdistribution of two introduced dung beetle species Digitonthophagus gazella (F.) and Euoniticellus intermedius (Reiche) (Coleoptera: Scarabaeidae) in an arid region of northern Mexico (Durango, Mexico)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {71},
	year = {1997},
	pages = {17-32},
	keywords = {aggregation, arid region, covariation, dung beetle, introduced species, spatial distribution},
	author = {Lobo, Jorge M. and Montes de Oca, Enrique}
}
@article {1976,
	title = {New data and observations on kleptoparasitic behaviour in dung beetles from temperate regions (Coleoptera: Scarabaeoidea)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {57},
	year = {1993},
	pages = {15-18},
	author = {Martin-Piera, Fermin and Lobo, Jorge M.}
}
@article {1672,
	title = {Informaciones ecologicas sobre Coprophanaeus (Coprophanaeus) telamon nevinsoni Arnaud \& Gamez (Coleoptera: Scarabaeinae: Phanaeini) en un sector de selva humeda submontana en Merida, Venezuela},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {22},
	number = {3},
	year = {2006},
	month = {Aug},
	pages = {95-105},
	keywords = {annual activity, carrion beetle, daily activity, habitat use, Scarabaeinae},
	author = {Gamez, Jorge and Mora, Enrique and Acconcia, Raffaele}
}
@article {2076,
	title = {Panamanian Scarabaeinae: taxonomy, distribution and habitats (Coleoptera, Scarabaeidae).},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {87},
	year = {2002},
	note = {digital copy},
	pages = {67-82},
	keywords = {behavior, dung beetle, life history mammals, Scarabaeinae, Sulcophanaeus},
	author = {Noriega, Jorge}
}
@article {2093,
	title = {Biogeography of the areas and Canthonini (Coleoptera: Scarabaeidae) of dry tropical foress in Mesoamerica and Colombia},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {23},
	number = {1},
	year = {2007},
	pages = {73-108},
	abstract = {This biogeographical analysis examines the historical, geological, climatic and ecological processes that have influenced the formation of the dry tropical forests (DTF) of Mesoamerica and Colombia, areas that are the setting for multiple biogeographical stories that in this case are illustrated by the patterns and evolutionary processes of Canthonini. In this study we test the hypothesis that the Canthonini fauna of dry tropical forests has a South American affinity. To this end, we compare extant species from a tract of dry tropical forest in Mexico, a second enclave in Costa Rica, four from the Caribbean region of Colombia and finally one from the north of Tolima in the Upper Magdalena River Valley, Colombia. The geomorphological characteristics of the enclaves of DTF are also compared, as are the geographical distribution and taxonomic affinities of each of the species found in these dry tropical forests. The biogeographical, historical and geological aspects of the enclaves were evaluated using a Parsimony Analysis of Endemicity (PAE), with two tropical rain forests as the outgroup: Leticia (Amazonas, Colombia) and Los Tuxtlas (Veracruz, Mexico). This study reveals the origin and distribution of Neotropical dry forests in the Pleistocene, and the establishment of its dry conditions during the Holocene. It also reveals apparent similarities among the Canthonini of the dry tropical forests of Mexico, Costa Rica and Colombia, with three geographical distribution patterns that correspond to different degrees of expansion towards the north and the diversification of evolutionary lines, and even species of South American origin. The comparison of the cladogram generated for species of Canthonini with that of the geological events that have occurred in the study regions indicates that the distribution of Canthonini in dry tropical forests began during the Pliocene with the re-establishment of the Panamanian connection, with no evidence of previous geomorphological events having any influence. On the other hand, there are few species shared with the tropical rain forests used as the outgroup for the cladograms. },
	keywords = {biogeographical patterns, Canthonini, Colombia, Costa Rica, dry tropical forest, Mexico, Scarabaeinae},
	author = {Padilla-Gil, Dora Nancy and Halffter, Gonzalo}
}
@article {2133,
	title = {Fauna de coleoptera melolonthidae y passalidae de Tzucacab y Conkal, Yucatan, Mexico},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {21},
	number = {2},
	year = {2005},
	pages = {15-49},
	abstract = {A study of the Melolonthidae and Passalidae beetle fauna of Conkal and Tzucacab, Yucatan, Mexico is presented. Data from 6,378 specimens were obtained during 2001-2002 with 74 days of collecting effort in tropical deciduous forest and secondary vegetation, located between 9-70 m of altitude. Forty nine species of the genera Phyllophaga, Diplotaxis, Anomala, Chasmodia, Macraspis, Plesiorutela, Pelidnota, Cyclocephala, Ligyrus, Strategus, Coelosis, Phileurus, Gymnetis, Cotinis, Hoplopyga, Balsameda, Amithao, Euphoria, Trigonopeltastes, Passalus, Heliscus and Verres, are recorded. Species were more abundant in April, May and June. Most of them were from Conkal, but higher species richness was found in Tzucacab. Eighteen species are reported for the first time from Yucatan State. Hoplopyga ocellata G. \& P. and Euphoria subguttata Burmeister are confirmed as part of the Mexican fauna. A key to the 49 species, as well as comparative data between scarab beetle fauna of Tzucacab, Conkal and other tropical localities of Mexico, are included. Some biogeographically, agricultural and conservational aspects of these fauna are briefly discussed.},
	keywords = {Amithao, Distribution, Euphoria, Hoplopyga, Melolonthidae, Passalidae, Phyllophaga, taxonomy, tropical deciduous forest.},
	author = {Reyes Novelo, Enrique and Moron, Miguel-Angel}
}
@article {2151,
	title = {Aspectos biologicos de Canthon lituratus (Germar, 1813) e Canthidium (Canthidium) megathopoides Boucomont, 1928 (Coleoptera, Scarabaeidae)},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {70},
	year = {1997},
	pages = {1-12},
	author = {Rodrigues, Sergio R. and Flechtmann, Carlos Alberto Hector}
}
@article {2262,
	title = {Consideracoes sobre forrageio arboreo por Scarabaeidae (Coleoptera, Scarabaeoidea), e dados sobre sua ocorrencia  em floresta tropical do Brasil},
	journal = {Acta Zoologica Mexicana (nueva serie)},
	volume = {72},
	year = {1997},
	pages = {55-61},
	keywords = {Atlantic rainforest, brazil, dung beetle, foraging, Scarabaeidae},
	author = {Vaz-de-Mello, Fernando Z. and Louzada, Julio N. C.}
}
