H.  Sato and Imamori, M., ?Nidification of an african ball-rolling scarab, Scarabaeus platynotus Baters (Coleoptera, Scarabaeidae)?, Kontyu, vol. 54, pp. 203-207, 1986.

H.  Sato and Imamori, M., ?Production of two brood pears from one dung ball in an african ball-roller, Scarabaeus aegyptiorum (Coleoptera, Scarabaeidae)?, Kontyu, vol. 54, pp. 381-385, 1986.