H. H.  Aschenborn, Loughnan, M. L., and Edwards, P. B., ?A simple assay to determine the nutrional suitability of cattle dung for coprophagous beetles ?, Entomologia experimentalis et applicata, vol. 53, pp. 73-79, 1989.

G. M.  Carpaneto, Mazziotta, A., and Piattella, E., ?Changes in food resources and conservation of scarab beetles: from sheep to dog dung in a green urban area of Rome (Coleoptera, Scarabaeoidea)?, Biological Conservation, vol. 123, pp. 547-556, 2005.

L.  Dormont, Epinat, G., and Lumaret, J. P., ?Trophic preferences mediated by olfactory cues in dung beetles colonizing cattle and horse dung?, Environmental Entomology, vol. 33, pp. 370-377, 2004.

A.  Goidanich and Malan, C. E., ?Sulla fonte di alimentazione e sulla microflora aerobica del nido pedotrofico e dell'apparato digerente delle larve di Scarabei coprofagi (Coleoptera Scarabaeidae)?, Atti Accad. Sc. Torino, vol. 96, pp. 575-628, 1962.

I.  Hanski, ?Nutritional ecology of dung and carrion feeding insects?, in Nutritional Ecology of Insects, Mites and Spiders, J. F.  Slanky and Rodriguez, J. G., Eds. New York: Wiley, 1987, pp. 837-884.

N.  Kadiri, Lobo, J. M., and Lumaret, J. P., ?Conséquences de l?interaction entre préférences pour l?habitat et quantite´ de ressources trophiques sur les communaute´s d?insectes coprophages (Coleoptera: Scarabaeoidea)?, Acta Oecologica-International Journal of Ecology, vol. 18, pp. 107-119, 1997.

I. Martinez M. and Vázquez, A., ?Influencia de algunos factores ambientales sobre la reproducciión en Canthon cyanellus cyanellus LeConte (Coleoptera: Scarabaeidae: Scarabdaeinae)?, Elytron, vol. 9, pp. 5-13, 1995.

M. I.  Martínez and Vázquez, A., ?Influencia de algunis factores ambientales sobre la reproducciión en Canthon cyanellus cyanellus LeConte (Coleoptera: Scarabaeidae: Scarabdaeinae)?, Elytron, vol. 9, pp. 5-13, 1995.

E. G.  Matthews, ?La biogeografía ecológia de los escarabajos del estiércol?, Acta Politécnica Mexicana, vol. 16, pp. 89-98, 1975.

E. G.  Matthews and Halffter, G., ?The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera, Scarabaeidae)?, Folia Entomologica Mexicana, vol. 12-14, pp. 1-312, 1966.

T.  Ochi and Kon, M., ?Dung beetles (Coleoptera, Scarabaeidae) utilizing the materials other than dung?, Nature and Insects, vol. 39, pp. 16-18, 2004.

T.  Schmitt, Krell, F. T., and Linsenmair, K. E., ?Quinone mixture as attractant for necrophagous dung beetles specialized on dead millipedes?, Journal of Chemical Ecology, vol. 30, pp. 731-740, 2004.

K.  Schoenly, ?Arthropods associated with bovine and equine dung in an ungrazedChihuahuan desert ecosystem?, Annals of the Entomological Society of America, vol. 76, pp. 790-796, 1983.

F. N.  Young and Goff, C. C., ?An annotated list of the arthropods found in burrows of the Florida Gopher Tortise (Gopherus polyphemus) (Daudin)?, Florida Entomologist, vol. 22, pp. 53-62, 1939.