S.  Caveney, Scholtz, C. H., and McIntyre, P., ?Patterns of daily flight activity in onitine dung beetles (Scarabaeinae: Onitini)?, Oecologia, vol. 103, pp. 444-452, 1995.

S. L.  Chown, Pistorius, P., and Scholtz, C. H., ?Morphological correlates of flightlessness in southern African Scarabaeinae (Coleoptera: Scarabaeidae): testing a condition of the water-conservation hypothesis?, Canadian Journal of Zoology, vol. 76, pp. 1123-1133, 1998.

M.  Dacke, Nordstrom, P., and Scholtz, C. H., ?Twilight orientation to polarised light in the crepuscular dung beetle Scarabaeus zambesianus?, The Journal of Experimental Biology, vol. 206, pp. 1535-1543, 2003.

M.  Dacke, Nordstrom, P., Scholtz, C. H., and Warrant, E. J., ?A specialized dorsal rim area for polarized light detection in the compound eye of the scarab beetle Pachysoma striatum?, Journal of Comp. Physiology A, vol. 188, pp. 211-216, 2002.

A. L. V.  Davis, Scholtz, C. H., and Deschodt, C., ?A dung beetle survey of selected Gauteng nature reserves: implications for conservation of the provincial scarabaeine fauna?, African Entomology, vol. 13, pp. 1-16, 2005.

A. L. V.  Davis, Scholtz, C. H., and Harrison, Jdu G., ?Cladistic, phenetic and biogeographical analysis of the flightless dung beetle genus, Gyronotus van Lansberge (Scarabaeidae: Scarabaeinae), in threatened eastern Afrotropical forests?, Journal of Natural History, vol. 35, pp. 1607-1625, 2001.

V. V.  Grebennikov, Ballerio, A., Ocampo, F. C., and Scholtz, C. H., ?Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae?, Systematic Entomology, vol. 29, pp. 524-543, 2004.

V. V.  Grebennikov and Scholtz, C. H., ?The basal phylogeny of Scarabaeoidea (Insecta: Coleoptera) inferred from larval morphology?, Invertebrate Systematics, vol. 18, pp. 321-348, 2004.

H. F.  Howden and Scholtz, C. H., ?Changes in a Texas USA Dung Beetle Community between 1975 and 1985 Coleoptera Scarabaeidae Scarabaeinae?, The Coleopterists Bulletin, vol. 40, pp. 313-316, 1986.

K.  Kruger, Lukhele, O. M., and Scholtz, C. H., ?Survival and reproduction of Euoniticellus intermedius (Coleoptera: Scarabaeidae) in dung following application of cypermethrin and flumethrin pour-ons to cattle?, Bulletin of Entomological Research, vol. 89, pp. 543-548, 1999.

K.  Kruger and Scholtz, C. H., ?Lethal and sublethal effects of ivermectin on the dung-breeding beetles Euoniticellus intermedius (Reiche) and Onitis alexis Klug (Coleoptera: Scarabaeidae)?, Agriculture, Ecosystems and Environment, vol. 61, pp. 123-131, 1997.

K.  Kruger and Scholtz, C. H., ?Possible risk posed by drift of the insect growth regulator pyriproxyfen to the rare dung beetle Circellium bacchus (F.) in a national park?, Journal of Insect Conservation, vol. 1, pp. 215-220, 1997.

K.  Kruger and Scholtz, C. H., ?Changes in the structure of dung insect communities after ivermectin usage in a grasslansd  ecosystem. II. Impact of ivermectin under high-rainfall conditions?, Acta Oecologica, vol. 19, pp. 439-451, 1998.

U.  Kryger, Cole, K. S., Tukker, R., and Scholtz, C. H., ?Biology and ecology of Circellium bacchus (Fabricius 1781) (Coleoptera, Scarabaeidae), a South African dung beetle of conservation concern?, Tropical Zoology, vol. 19, pp. 185-207, 2006.

U.  Kryger, Deschodt, C., Davis, A. L., and Scholtz, C. H., ?Effects of cattle treatment with a cypermethrin/cymiazol spray on survival and reproduction of the dung beetle species Euoniticellus intermedius (Coleoptera: Scarabaeidae)?, Bulletin of Entomological Research, vol. 96, pp. 597-603, 2006.

U.  Kryger, Deschodt, C., Davis, A. L. V., and Scholtz, C. H., ?Effects of cattle treatment with a fluazuron pour-on on survival and reproduction of the dung beetle species Onthophagus gazella (Fabricius)?, Veterinary Parasitology, vol. 143, pp. 380-384, 2007.

E.  Le Roux, Scholtz, C. H., Kinahan, A. A., and Bateman, P. W., ?Pre- and post-copulatory mate selection mechanisms in an african dung beetle, Circellium bacchus (Coleoptera: Scarabaeidae)?, Journal of Insect Behavior, vol. 21, pp. 111-122, 2008.

P.  Holter and Scholtz, C. H., ?Are ball-rolling (Scarabaeini, Gymnopleurini, Sisyphini) and tunnelling scarabaeine dung beetles equally choosy about the size of ingested dung particles??, Ecological Entomology, vol. 30, pp. 700-705, 2005.

P.  Holter and Scholtz, C. H., ?What do dung beetles eat??, Ecological Entomology, vol. 32, pp. 690-697, 2007.

P.  Holter, Scholtz, C. H., and Wardhaugh, K. G., ?Dung feeding in adult scarabaeines (tunnellers and endocoprids): even large dung beetles eat small particles.?, Ecological Entomology, vol. 27, pp. 169-176, 2002.

K.  T. Philips, Pretorius, E., and Scholtz, C. H., ?A phylogenetic analysis of dung beetles (Scarabaeinae: Scarabaeidae): unrolling an evolutionary history?, Invertebrate Systematics, vol. 18, pp. 53-88, 2004.

R.  Pretorius, T. Philips, K., and Scholtz, C. H., ?Geometric morphometrics, the metendosternite and its use in phylogenetics of the Scarabaeinae (Coleoptera)?, Elytron, vol. 14, pp. 125-148, 2000.

C. H.  Scholtz, Harrison, Jdu G., and Grebennikov, V. V., ?Dung beetle (Scarabaeus (Pachysoma)) biology and immature stages: reversal to ancestral states under deset conditions (Coleoptera: Scarabaeidae)??, Biological Journal of the Linnean Society, vol. 83, pp. 453-460, 2004.

C. L.  Sole, Scholtz, C. H., and Bastos, A. D. S., ?Phylogeography of the Namib Desert dung beetles Scarabaeus (Pachysoma) MacLeay (Coleoptera: Scarabaeidae)?, Journal of Biogeography, vol. 32, pp. 75-84, 2005.

P.  B. Tshikae, Davis, A. L. V., and Scholtz, C. H., ?Trophic associations of a dung beetle assemblage (Scarabaeidae: Scarabaeinae) in a woodland savanna of Botswana?, Environmental Entomology, vol. 37, pp. 431-441, 2008.