A. L. V.  Davis, ?Annual age structure patterns in Afrotropical dung beetles (Coleoptera: Scarabaeidae) under winter rainfall climate?, Journal of African Zoology, vol. 107, pp. 397-411, 1993.

A. L. V.  Davis, ?Community organization in a South African, winter rainfall, dung beetle assemblage (Coleoptera: Scarabaeidae)?, Acta Oecologica, vol. 15, pp. 727-738, 1994.

A. L. V.  Davis, ?Compositional differences between dung beetle (Coleoptera: Scarabaeidae s. str.) assemblages in winter and summer rainfall climates?, African Entomology, vol. 2, pp. 45-51, 1994.