A. L. V.  Davis, ?Biogeographical groups in a southern African, winter rainfall, dung beetle assemblage (Coleoptera: Scarabaeidae) - consequences of climatic history and habitat fragmentation?, African Journal of Ecology, vol. 31, pp. 306-327, 1993.

A. L. V.  Davis, ?Daily weather variation and temporal dynamics in an Afrotropical dung beetle community (Coleoptera: Scarabaeidae)?, Acta Oecologica, vol. 16, pp. 641-656, 1995.

A. L. V.  Davis and Dewhurst, C. F., ?Climatic and biogeographical associations of Kenyan and northern Tanzanian dung beetles (Coleoptera: Scarabaeidae)?, African Journal of Ecology, vol. 31, pp. 290-305, 1993.

A. A.  Kirk, ?Dung beetles (Coleoptera: Scarabaeidae) active in patchy forest and pasture habitats in Santa Cruz Province, Bolivia, during spring?, Folia Entomologica Mexicana, vol. 84, pp. 45-54, 1992.

M.  Palmer, ?Testing for seasonal displacement in a dung beetle guild?, Ecography, vol. 18, pp. 173-177, 1995.