F.  Ervik and Knudsen, J. T., ?Water lilies and scarabs: faithful partners for 100 million years??, Biological Journal of the Linnean Society, vol. 80, pp. 539-543, 2003.

A. P.  Moczek, Bruhl, C., and Krell, F. - T., ?Linear and threshold-dependent expression of secondary sexual traits in the same individual: insights from a horned beetle (Coleoptera: Scarabaeidae)?, Biological Journal of the Linnean Society, vol. 83, pp. 473-480, 2004.

C. H.  Scholtz, Harrison, Jdu G., and Grebennikov, V. V., ?Dung beetle (Scarabaeus (Pachysoma)) biology and immature stages: reversal to ancestral states under deset conditions (Coleoptera: Scarabaeidae)??, Biological Journal of the Linnean Society, vol. 83, pp. 453-460, 2004.