F. G.  Horgan, ?Aggregation and coexistence of dung beetles in montane rain forest in deforested siters in central Peru?, Journal of Tropical Ecology, vol. 22, pp. 359-370, 2006.

F. G.  Horgan, ?Dung beetle assemblages in forests and pastures of El Salvador: a functional comparison?, Biodiversity Conservation, vol. 17, pp. 2961-2978, 2008.