J.  Hunt, Simmons, L. W., and Kotiaho, J. S., ?A cost of maternal care in the dung beetle Onthophagus taurus??, Journal of Evolutionary Biology, vol. 15, pp. 57-64, 2002.

J. S.  Kotiaho, ?Sexual selection and condition dependence of courtship display in three species of horned dung beetles?, Behavioral Ecology, vol. 13, pp. 791-799, 2002.

J. S.  Kotiaho, Simmons, L. W., and Tomkins, J. L., ?Towards a resolution of the lek paradox?, Nature, vol. 410, pp. 684-686, 2001.