D. J.  Emlen, ?A new method for observing underground and soil surface behaviors?, The Coleopterists Bulletin, vol. 47, pp. 243-244, 1993.

D. J.  Emlen, ?Alternative reproductive tactics and male-dimorphism in the horne beetle Onthophagus acuminatus (Coleoptera: Scarabaeidae)?, Behavioral Ecology and Sociobiology, vol. 41, pp. 335-341, 1997.

D. J.  Emlen, ?Diet alters male horn allometry in the beetle Onthophagus acuminatus (Coleoptera: Scarabaeidae)?, Proceedings of the Royal society of London B, vol. 264, pp. 567-574, 1997.

D. J.  Emlen, ?Integrating development with evolution: a case study with beetle horns?, Bioscience, vol. 50, pp. 403-418, 2000.

D. J.  Emlen, ?Costs and the diversification of exaggerated animal structures?, Science, vol. 291, pp. 1534-1536, 2001.

D. J.  Emlen, Marangelo, J., Ball, B., and Cunningham, C., ?Diversity in the weapons of sexual selection: horn evolution in the beetle genus Onthophagus (Coleoptera: Scarabaeidae)?, Evolution, vol. 59, pp. 1060-1084, 2005.

D. J.  Emlen and H. Nijhout, F., ?Hormonal control of male horn lenght dimorphism in the dung beetle Onthophagus taurus (Coleoptera: Scarabaeidae)?, Journal of Insect Physiology, vol. 45, pp. 45-53, 1999.

D. J.  Emlen and H. Nijhout, F., ?The development and evolution of exaggerated morphologies in insects?, Annual Review of Entomology, vol. 45, pp. 661-708, 2000.

D. J.  Emlen and T. Philips, K., ?Phylogenetic evidence for an association between tunneling behavior and the evolution of horns in dung beetles (Coleoptera : Scarabaeidae : Scarabaeinae)?, The Coleopterists Bulletin, vol. 60, pp. 47-56, 2006.

A. P.  Moczek and Emlen, D. J., ?Male horn dimorphism in the scarab beetle, Onthophagus taurus: do alternative reproductive tactics favour alternative phenotypes?, Animal Behavior, vol. 59, pp. 459-466, 2000.

A. P.  Moczek, Hunt, J., Emlen, D. J., and Simmons, L. W., ?Threshold evolution in exotic populations of a polyphenic beetle?, Evolutionary Ecology Research, vol. 4, pp. 587-601, 2002.