A. J.  Davis, Huijbregts, H., and Krikken, J., ?The role of local and regional processes in shaping dung beetle communities in tropical forest plantations in Borneo?, Global Ecology and Biogeography, vol. 9, pp. 281-292, 2000.

J. N. C.  Louzada, ?Considerations on the perching behavior of tropical dung beetles (coleoptera: Scarabaeidae)?, Revista Brasileira de Entomologia, vol. 41, pp. 125-128, 1998.

S.  Spector and Ayzama, S., ?Rapid turnover and edge effects in dung beetle assemblages (Scarabaeidae) at a Bolivian Neotropical forest-savanna ecotone?, Biotropica, vol. 35, pp. 394-404, 2003.