L.  Brussaard and Visser, W. J. F., ?Dung Exploitation of the Dung Beetle Typhaeus-Typhoeus Coleoptera Geotrupidae?, Oecologia Berlin, vol. 72, pp. 21-27, 1987.

A. L. V.  Davis, ?Geographical Distribution of Dung Beetles (Coleoptera: Scarabaeidae) and Their Seasonal Activity in Southwestern Cape Province South Africa?, Journal of the Entomological Society of Southern Africa, vol. 50, pp. 275-285, 1987.

I.  Hanski and Kuusela, S., ?Dung Beetle Communities in the Aland Archipelago Finland?, Acta Entomologica Fennica, vol. 42, pp. 36-42, 1983.

H. F.  Howden, Howden, A. T., and Storey, R. I., ?Nocturnal Perching of Scarabaeine Dung Beetles Coleoptera Scarabaeidae in an Australian Tropical Rain Forest?, Biotropica, vol. 23, pp. 51-57, 1991.

H. F.  Howden and Scholtz, C. H., ?Changes in a Texas USA Dung Beetle Community between 1975 and 1985 Coleoptera Scarabaeidae Scarabaeinae?, The Coleopterists Bulletin, vol. 40, pp. 313-316, 1986.

J.  Hunt, Kotiaho, J. S., and Tomkins, J. L., ?Dung pad residence time covaries with male morphology in the dung beetle Onthophagus taurus?, Ecological Entomology, vol. 24, pp. 174-180, 1999.

J.  Hunt and Simmons, L. W., ?Patterns of parental provisioning covary with male morphology in a horned beetle (Onthophagus taurus) (Coleoptera: Scarabaeidae)?, Behavioral Ecology and Sociobiology, vol. 42, pp. 447-451, 1998.

J.  Hunt and Simmons, L. W., ?Maternal and paternal effects on offspring phenotype in the dung beetle Onthophagus taurus?, Evolution, vol. 54, pp. 936-941, 2000.

J. M.  Lobo, ?Modification in the dung beetles communities on highland grasslands of the Iberian Central System (Spain) along an altitudinal gradient?, Acta Zoologica Mexicana Nueva Serie, pp. 15-31, 1992.

J. M.  Lobo, ?Diversity, biogeographical considerations and spatial structure of a recently invaded dung beetle (Coleoptera: Scarabaeoidea) community in the Chihuahuan desert?, Global Ecology and Biogeography Letters, vol. 5, pp. 342-352, 1996.

J. M.  Lobo, ?Species diversity and composition of dung beetle (Coleoptera: Scarabaeoidea) assemblages in North America?, Canadian Entomologist, vol. 132, pp. 307-321, 2000.

J. M.  Lobo, ?Decline of roller dung beetle (Scarabaeinae) populations in the Iberian peninsula during the 20th century?, Biological Conservation, vol. 97, pp. 43-50, 2001.

J. M.  Lobo and Davis, A. L. V., ?An intercontinental comparison of dung beetle diversity between two mediterranean-climatic regions: Local versus regional and historical influences?, Diversity and Distributions, vol. 5, pp. 91-103, 1999.

J. M.  Lobo, Sanmartin, I., and Martin, P. Fermin, ?Diversity and spatial turnover of dung beetle (Coleoptera: Scarabaeoidae) communities in a protected area of South Europe (Donana National Park, Huelva, Spain)?, Elytron, vol. 11, pp. 71-88, 1997.

J. P.  Lumaret, Kadiri, N., and Bertrand, M., ?Changes in Resources Consequences for the Dynamics of Dung Beetle Communities?, Journal of Applied Ecology, vol. 29, pp. 349-356, 1992.

A.  Estrada and Coates-Estrada, R., ?Howler Monkeys Alouatta-Palliata Dung Beetles Scarabaeidae and Seed Dispersal Ecological Interactions in the Tropical Rain Forest of Los Tuxtlas Mexico?, Journal of Tropical Ecology, vol. 7, pp. 459-474, 1991.

M. E.  Favila and Halffter, G., ?The use of indicator groups for measuring biodiversity as related to community structure and function?, Acta Zoologica Mexicana Nueva Serie, pp. 1-25, 1997.

E.  Galante, Garcia, R. M., Barrera, I., and Galindo, P., ?Comparison of Spatial Distribution Patterns of Dung-Feeding Scarabs Coleoptera Scarabaeidae Geotrupidae in Wooded and Open Pastureland in the Mediterranean Dehesa Area of the Iberian Peninsula?, Environmental Entomology, vol. 20, pp. 90-97, 1991.

A. P.  Moczek and Emlen, D. J., ?Proximate determination of male horn dimorphism in the beetle Onthophagus taurus (Coleoptera: Scarabaeidae)?, Journal of Evolutionary Biology, vol. 12, pp. 27-37, 1999.

E.  Morelli, Gonzalez, V. Patricia, and Canziani, C., ?Nesting and life-cycle of Sulcophanaeus menelas (Laporte, 1840) (Coleoptera, Scarabaeidae)?, Elytron Barcelona, vol. 10, pp. 11-22, 1996.

V. G.  Nealis, ?Habitat Associations and Community Analysis of South Texas Dung Beetles Coleoptera Scarabaeinae?, Canadian Journal of Zoology, vol. 55, pp. 138-147, 1977.

C.  Rojewski, ?The Role of Dung Beetles in Nature and in the Economy?, Przeglad Zoologiczny, vol. 24, pp. 431-438, 1980.

H.  Sato, ?Male participation in nest building in the dung beetle Scarabaeus catenatus (Coleoptera: Scarabaeidae): Mating effort versus paternal effort?, Journal of Insect Behavior., vol. 11, pp. 833-843, 1998.

K.  Vulinec, ?Iridescent dung beetles: A different angle?, Florida Entomologist, vol. 80, pp. 132-141, 1997.