J. R.  Anderson, Merrit, R. W., and Loomis, E. C., ?The insect-free cattle dropping and its relationship to increased dung fouling of rangeland pastures.?, Journal of Economic Entomology, vol. 77, pp. 133-141, 1984.

G. C.  Walsh, Cordo, H. A., Briano, J. A., Gandolfo, D. E., and Logarzo, G. A., ?Laboratory culture of beneficial dung scarabs?, Journal of Economic Entomology, vol. 90, pp. 124-129, 1997.