H. H.  Aschenborn, Loughnan, M. L., and Edwards, P. B., ?A simple assay to determine the nutrional suitability of cattle dung for coprophagous beetles ?, Entomologia experimentalis et applicata, vol. 53, pp. 73-79, 1989.

T. J.  Ridsdill-Smith, Hayles, L., and Palmer, M. L., ?Competition between the bush fly and dung beetles in dung of differing characteristics?, Entomologia Experimentalis et Applicata, vol. 41, pp. 83-90, 1986.