A. L. V.  Davis, ?Residence and breeding of Oniticellus (Coleoptera: Scarabaeidae) within cattle pads: inhibition by dung-burying beetles?, Journal of the Entomological Society of Southern Africa, vol. 52, pp. 229-236, 1989.

T. J.  Ridsdill-Smith and Hayles, L., ? Re-examination of competition between Musca vetustissima Walker (diptera: Muscidae) larvae and seasonal changes in favorability of catle dung.?, Journal of the Australian Entomological Society, vol. 28, pp. 105-111, 1989.

K.  Yokoyama, Kai, H., Koga, T., and Kawaguchi, S., ?Effect of dung beetle Onthophagus lenzii H. on nitrogen transformation in cow dung and dung balls?, Soil Science and Plant Nutrition, vol. 37, pp. 341-345, 1989.