S.  Caveney, Scholtz, C. H., and McIntyre, P., ?Patterns of daily flight activity in onitine dung beetles (Scarabaeinae: Onitini)?, Oecologia, vol. 103, pp. 444-452, 1995.

P.  McIntyre and Caveney, S., ?Superposition optics and the time of flight in onitine dung beetles?, Journal of Comp. Physiology A, vol. 183, pp. 45-60, 1998.