Y.  Hingrat and Feer, F., ?Effects of forest fragmentation on the activity of dung beetles?, Revue D Ecologie-La Terre Et La Vie, pp. 165-179, 2002.

N.  Kanda, Yokota, T., Shibata, E., and Sato, H., ?Diversity of dung-beetle community in declining Japanese subalpine forest caused by an increasing sika deer population?, Ecological Research, vol. 20, pp. 135-141, 2005.

K.  Feeley, ?The role of clumped defecation in the spatial distribution of soil nutrients and the availability of nutrients for plant uptake?, Journal of Tropical Ecology, vol. 21, pp. 99-102, 2005.

F.  Feer, ?Dung and carrion beetles of the rain forest of French Guiana: composition and structure of the guild?, Annales De La Societe Entomologique De France, vol. 36, pp. 29-43, 2000.

G.  Ponce-Santizo, Andresen, E., Cano, E., and Cuaron, A. D., ?Primary dispersal of seeds by primates and secondary dispersal by dung beetles in Tikal, Guatemala?, Biotropica, vol. 38, pp. 390-397, 2006.

P. R.  Stevenson, Castellanos, M. C., Pizarro, J. C., and Garavito, M., ?Effects of seed dispersal by three ateline monkey species on seed germination at Tinigua National Park, Colombia?, International Journal of Primatology, vol. 23, pp. 1187-1204, 2002.