J. L.  Dew, ?Seed dispersal by two amazonian primates, with notes on dung beetles?, American Zoologist, vol. 39, 1999.

F.  Martin-Piera and Lobo, J. M., ?A comparative discussion of trophic preferences in dung beetle communities?, Miscellania Zoologica Barcelona, vol. 19, pp. 13-31, 1996.