K.  Takigawa, Sato, M., Yanagi, A., Asaoka, S., Yamashita, N., Nakanishi, Y., Manda, M., Yanagita, K., and Hayakawa, H., ?Activity of dung beetle and its effect on dung patch existence in grazed pasture?, Grassland Science, vol. 43, 1997.

D.  Yamada, Imura, O., Shi, K., and Shibuya, T., ?Effect of tunneler dung beetles on cattle dung decomposition, soil nutrients and herbage growth ?, Grassland Science, vol. 53, pp. 121-129, 2007.