M.  Bertone, ?Dung beetles (Coleoptera: Scarabaeidae and Geotrupidae) in North Carolina Pasture Ecosystem?, North Carolina University, Raleigh, 2006.

M.  Bertone, Green, J., Washburn, S., Poore, M., Sorenson, C., and D. Watson, W., ?Seasonal activity and species composition of dung beetles (Coleoptera : Scarabaeidae and Geotrupidae) inhabiting cattle pastures in North Carolina?, Annals Of The Entomological Society Of America, vol. 98, pp. 309-321, 2005.

M. A.  Bertone, ?Dung beetles (Coleoptera: Scarabaeidae and Geotrupidae) of North Carolina Cattle Pastures  and their Implications for Pasture Improvement?, North Carolina University, Raleigh, 2004.

M.  Wallace and Richardson, R. H., ?Observations of urban dung beetles utilizing dog feces (Coleoptera: Scarabaeidae)?, The Coleopterists Bulletin, vol. 59, pp. 400-401, 2005.