B. V.  Burger, Petersen, W. G. B., Weber, W. G., and Munro, Z. M., ?Semiochemicals of the Scarabaeinae. VII: Identification and synthesis of EAD-active constituents of abdominal sex attracting secretion of the male dung beetle, Kheper subaeneus?, Journal of Chemical Ecology, vol. 28, pp. 2527-2539, 2002.