A. L. V.  Davis and Scholtz, C. H., ?Local and regional species ranges of a dung beetle assemblage from the semi-arid Karoo/Kalahari margins, South Africa?, Journal of Arid Environments, vol. 57, pp. 61-85, 2004.

S.  F. Pinero and Avila, J. M., ?Dung-insect community composition in arid zones of south-eastern Spain?, Journal of Arid Environments, vol. 56, pp. 303-327, 2004.