A. J.  Davis and Lantoh, S., ?Dung beetles collected from Rafflesia pricei Meijer at the Tambunan Reserve, Sabah, Malaysia?, Malayan Nature Journal , vol. 50, pp. 39-41, 1996.

F.  Ervik and Knudsen, J. T., ?Water lilies and scarabs: faithful partners for 100 million years??, Biological Journal of the Linnean Society, vol. 80, pp. 539-543, 2003.

S.  Sakai and Inoue, T., ?A new pollination system: Dung-beetle pollination discovered in Orchidantha inouei (Lowiaceae, Zingiberales) in Sarawak, Malaysia?, American Journal of Botany, vol. 86, pp. 56-61, 1999.