M. Clara Castellanos, S., F. Escobar, and Stevenson, P. R., ?Dung beetles (Scarabaeidae: Scarabaeinae) attracted to woolly monkey (Lagothrix lagothricha Humboldt) dung at Tinigua National Park, Colombia?, The Coleopterists Bulletin, vol. 53, pp. 155-159, 1999.

A. J.  Davis, ?Perching behaviour in Bornean dung beetles (Coleoptera : Scarabaeidae)?, The Coleopterists Bulletin, vol. 53, pp. 365-370, 1999.

P.  Gonzalez-Vainer and Morelli, E., ?Phenology and biology of the Dung beetle Onthophagus hirculus Mannerheim (Coleoptera : Scarabaeidae)?, The Coleopterists Bulletin, vol. 53, pp. 303-309, 1999.

F. - T.  Krell, ?Dung beetles unharmed by army ants in tropical Africa (Coleoptera: Scarabaeidae; Hymanoptera: Formicidae, Dorylinae)?, The Coleopterists Bulletin, vol. 53, pp. 325-328, 1999.

I. Martinez M. and R., M. Cruz, ?The effects of male glandular secretions on female endocrine centers in Canthon cyanellus cyanellus LeConte (Coleoptera: Scarabaeidae: Scarabaeinae)?, The Coleopterists Bulletin, vol. 53, pp. 208-216, 1999.

B. C.  Ratcliffe and Smith, A. B. T., ?New species of Canthonella Chapin (Scarabaeidae: Scarabaeinae) from Amazonian Brasil?, The Coleopterists Bulletin, vol. 53, pp. 1-7, 1999.