R. D.  Hughes, ?Introduced Dung Beetles and Australian Pasture Ecosystems - Papers Presented at a Symposium During Meeting of Australia- and-New-Zealand-Association-for-Advancement-of-Science at Canberra in January 1975?, Journal of Applied Ecology, vol. 12, pp. 819-837, 1975.

R. D.  Hughes, Tyndale-Biscoe, M., and Walker, J., ?Effects of introduced dung beetles (Coleoptera: Scarabaeinae) on the breeding and abundance of the Australian bushfly Musca vetustissima Walker (Diptera: Muscidae).?, Bulletin of Entomological Research, vol. 68, pp. 361-372, 1978.

J.  Feehan, Hughes, R. D., Bryce, M. A., and Runko, S., ?Bush fly abundance and population events in relation to dung beetles catches on the south coast of New South Wales?, Journal of the Australian Entomological Society, vol. 24, pp.  37-43, 1985.