R.  Madewell and Moczek, A. P., ?Horn possession reduces maneuverability in the horn-polyphenic beetle, Onthophagus nigriventris?, Journal of Insect Science, vol. 6, pp. 1-10, 2006.

A. P.  Moczek, ?Pupal remodeling and the development  and evolution of sexual dimorphism in horned beetles?, The American Naturalist, vol. 168, pp. 711-729, 2006.