L.  Brussaard and Visser, W. J. F., ?Dung Exploitation of the Dung Beetle Typhaeus-Typhoeus Coleoptera Geotrupidae?, Oecologia Berlin, vol. 72, pp. 21-27, 1987.

A. L. V.  Davis, ?Geographical Distribution of Dung Beetles (Coleoptera: Scarabaeidae) and Their Seasonal Activity in Southwestern Cape Province South Africa?, Journal of the Entomological Society of Southern Africa, vol. 50, pp. 275-285, 1987.