A. L. V.  Davis, ?Habitat fragmentation in southern Africa and distributional response patterns in five specialist or generalist dung beetle families (Coleoptera)?, African Journal of Ecology, vol. 32, pp. 192-207, 1994.

P. S.  Giller and Doube, B. M., ?Spatial and temporal co-occurrence of competitors in Southern African dung beetle communities?, Journal of Animal Ecology, vol. 63, pp. 629-643, 1994.

J. M.  Lobo and Montes, D. Oca Enriqu, ?Local distribution and coexistence of Digitonthophagus gazella (Fabricius, 1787) and Onthophagus batesi Howden and Cartwright 1963 (Coleoptera: Scarabaeidae)?, Elytron, vol. 8, pp. 117-127, 1994.